In: Red Wine and Health ISBN 978-1-60692-718-2
Editor: Paul O’Byrne © 2009 Nova Science Publishers, Inc.
Chapter 6
BIOGENIC AMINES IN WINES: A REVIEW
Fernanda Galgano, Marisa Caruso and Fabio Favati
Dept. of Biology, DBAF, University of Basilicata
Viale dell’Ateneo Lucano, 10, 85100 – Potenza - Italy
ABSTRACT
Biogenic amines are basic nitrogen compounds that can have toxicological effects on
human health when present in foods and beverages at significant levels. These
compounds are produced in small amounts in all living organisms and play an important
role in cell growth and development, as well as in protecting stressed cells. Their
presence at dangerous levels is principally attributed to microbial decarboxylation of the
corresponding amino acids, and hence the occurrence of biogenic amines can be related
to both food safety and food spoilage, becoming a useful index for the assessment of food
quality and of the related manufacturing practices.
Nowadays an increasing attention is given to the presence in wine of biogenic
amines, whose level and the possible synergic effect with ethanol may represent a
significant risk for some consumers. The major toxicological implications are related to
aromatic amines, such as histamine, tyramine, and 2-phenylethylamine; nevertheless, for
these amines it is quite difficult to establish the exact threshold of toxicity, which
depends on the efficiency of the detoxification mechanisms of different individuals.
However, some countries have set limits for histamine in wine ranging from 2 to 10
mg/L, while for tyramine levels exceeding 10 mg/L in beverages should be considered
unsafe. Several other amines may be found in wine, among them polyamines, that can
enhance the adverse effect of aromatic amines, or cause negative consequences on wine
aroma. Moreover, spermine and spermidine, that have secondary group, are involved in
nitrosamine formation, compounds with a known cancerous action. Volatile amines, such
as methylamine and ethylamine, come from amination of non-nitrogen compounds, such
as aldehydes and ketones, and they have not a toxic action, but can exert a negative effect
on wine aroma.
Biogenic amines usually found in wine are cadaverine, histamine, 2-
phenylethylamine, putrescine and tyramine; agmatine and ethanolamine can be abundant,
but they are generally little investigated. Low biogenic amine amounts, as normal
constituents of the raw materials, can be released in must from grape and pulp during the
winemaking process, and the biogenic amine concentration may increase as consequence
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