Psychopharmacology(1994) 114: 123-130
Psychopharmacology
© Springer-Verlag 1994
Atypical antipsychotics, clozapine and sulpiride do not antagonise
amphetamine-induced stereotyped locomotion
Susan Moore, Paul Kenyon
Department of Psychology,University of Plymouth, Drake Circus, Plymouth PL4 8AA, United Kingdom
Received: 18 November 1992 / Final version: 20 July 1993
Abstract. An automated tracking system which converted
an animal's path between quadrants of a circular open
field into a series of trips was used to analyse stereotyped
locomotion in amphetamine treated rats. Amphetamine
(3.5 mg/kg) increased the horizontal distance moved and
the number and proportion of thigmotaxic trips around
the perimeter of the apparatus (length 4 trips). To investi-
gate the hypothesis that classic antipsychotics, but not
atypical antipsychotics, would antagonise the repetitive
boundary patrolling associated with amphetamine-in-
duced hyperactivity, animals were pretreated with
haloperidol (0.01, 0.025, 0.05, 0.075 mg/kg), clozapine (5,
10, 20 mg/kg) or ( +__ )sulpiride (10, 20, 50 mg/kg) 30 min
before 3.5 mg/kg amphetamine. The results showed that
the classic antipsychotic haloperidol antagonised both
hyperactivity and the increased proportion of length
4 trips. In marked contrast, the atypical antipsychotics
clozapine and sulpiride antagonised hyperactivity but did
not reduce the proportion of length 4 trips. The inability
of atypical antipsychotics to reduce the repetitive bound-
ary patrolling associated with amphetamine-induced hy-
peractivity is consistent with the action of these drugs on
other forms of amphetamine-induced stereotyped behav-
iour, and indicates that locomotor routes under ampheta-
mine are stereotyped. The measurement of trip lengths
provides a sensitive tool for examining drug action on the
spatial distribution of open field locomotion.
Key words: Amphetamine - Locomotor patterns
- Stereotyped locomotion - Haloperidol - Clozapine
- Sulpiride
Antagonism of amphetamine-induced behaviours in ani-
mals has been used to study the underlying mechanisms of
antipsychotic drug treatment, and to detect novel anti-
psychotic agents. Considerable attention has focused on
amphetamine-induced stereotyped behaviours such as
sniffing, licking and gnawing because these behaviours are
reliably antagonised by classic antipsychotics such as
haloperidol and chlorpromazine (Szechtman et al. 1988).
Correspondence to: S Moore
However, it is now clear that a group of clinically effective
drugs-so-called atypical antipsychotics-do not antag-
onise all the components of the amphetamine-induced
syndrome.
Tschanz and Rebec (1989) reported that atypical anti-
psychotics vary in their ability to antagonise specific am-
phetamine-induced behaviours. For example, clozapine
blocks sniffing but not the rearing or head bobbing pro-
duced by the drug. Paradoxically, several researchers have
reported that clozapine and other atypical antipsychotics
actually enhance the stereotyped head movements and
sniffing induced by 2-5 mg/kg amphetamine (Robertson
and MacDonald 1985; Sharp et al. 1986).
The conflicting reports of the effects of atypical anti-
psychotics on amphetamine-induced stereotyped behavi-
ours, and the observation that atypical antipsychotics are
less likely to produce Parkinson-like extrapyramidal side
effects which are associated with classic antipsychotic
drugs, has cast doubt on the utility of amphetamine antag-
onism as a screening technique for antipsychotic potential
(Robertson and MacDonald 1985; Tschanz and Rebec
1989), and led to the suggestion that antagonism of am-
phetamine-induced stereotypy predicts a compound's po-
tential to induce unwanted extrapyramidal side effects in
humans (Ljungberg and Ungerstedt 1985). However, clas-
sic and atypical antipsychotics share the ability to antag-
onise amphetamine-induced hyperactivity, and this may
form the basis of a more appropriate test for antipsychotic
potential (Rebec and Bashore 1984; Ljungberg and Un-
gerstedt 1985).
At low doses, amphetamine increases open-field loco-
motion; with increasing dose the route taken by an animal
in the open field becomes increasingly repetitive (Schior-
ring 1979; Mueller et al. 1989a; Geyer 1990). Because of
this, several authors have argued that locomotion exhib-
ited under higher (3-5 mg/kg) doses of amphetamine is
stereotyped. We have recently developed a technique for
measuring the spatial distribution of open-field loco-
motion (Kenyon 1990, 1991; Kenyon et al. 1992), based on
the approach described by Mueller et al. (1989a, b). The
subject's sequence of movements between adjacent quad-
rants of the apparatus is subdivided into a series of "trip
lengths" A trip is determined by the extent of forward
locomotion without change of direction. A change of