Inland Waters (2014) 4, pp. 233-242
© International Society of Limnology 2014
DOI: 10.5268/IW-4.2.700
233
Article
Assigning hydrogen, carbon, and nitrogen isotope values for
phytoplankton and terrestrial detritus in aquatic food web studies
Carol Yang
1
*, Grace M. Wilkinson
1
, Jonathan J. Cole
2
, Stephen A. Macko
1
, and Michael L. Pace
1
1
University of Virginia, Charlottesville, VA USA
2
Cary Institute of Ecosystem Studies, Millbrook, NY USA
* Corresponding author: cy5n@virginia.edu
Received 30 September 2013; accepted 22 January 2014; published 29 April 2014
Abstract
Studies designed to assess the resources supporting aquatic consumers using stable isotope analysis require measure-
ments of the potential end members (basal resources). While some basal resources are easily measured, it is often
difficult to physically separate phytoplankton (one potential end member) from other components in seston. Further,
terrestrial materials entering aquatic ecosystems undergo diagenetic change, potentially altering isotope composition
and making it difficult to assign end member values. We tested techniques for determining the isotopic hydrogen (δ
2
H),
carbon (δ
13
C), and nitrogen (δ
15
N) values of terrestrial and phytoplankton end members in seston. Long term in situ leaf
decomposition experiments were performed. No appreciable change was found in the isotope values of degraded
material (mean change 3.6‰ for δ
2
H, 0.0‰ for δ
13
C, and −0.1‰ for δ
15
N). We conclude that the isotope values of
terrestrial plant material can be used to assign end members for terrestrial detritus. Using samples collected from 10
lakes with phytoplankton-dominated seston, we compared 3 published methods for estimating the δ
13
C and δ
15
N of
phytoplankton. One method, which corrected bulk particulate organic matter (POM) isotope values based on a δ
2
H
mixing model, accurately predicted measured phytoplankton δ
13
C. Another method, which used a C:N mixing model
to correct bulk POM, also performed well. A new method, proposed here, modified seston isotope values using the
difference in C:N of phytoplankton and terrestrial material in a δ
2
H mixing model and correctly predicted measured
phytoplankton δ
15
N. We recommend estimating phytoplankton δ
13
C and δ
15
N by correcting bulk POM using a δ
2
H
mixing model, with the C:N modification proposed here for δ
15
N.
Key words: food web, mixing models, phytoplankton, stable isotope analysis, terrestrial organic matter
Introduction
Stable isotope analysis is a common tool for evaluating
resource availability, trophic structure, and consumer
basal resource use. In aquatic ecosystems, stable isotope
values of carbon (
13
C/
12
C: δ
13
C), nitrogen (
15
N/
14
N: δ
15
N),
and hydrogen (
2
H/
1
H: δ
2
H) can be employed to quantify
the use of terrestrial (allochthonous) organic matter by
aquatic consumers (Marcarelli et al. 2011). Methodologi-
cal difficulties arise when quantifying consumer basal
resources using mixing models, however, because
assigning appropriate end member values for some source
materials is complicated. For example, phytoplankton are
often difficult to physically separate from bulk seston
(Hamilton et al. 2005). In addition, some materials with
long residence times (e.g., pelagic and benthic detritus)
may undergo changes in isotope ratios over time.
Although H isotopes may present an advantage over
C and N isotopes owing to the large end member
separation (Doucett et al. 2007), uncertainties remain in
estimating the incorporation of environmental water in
consumer tissues (Solomon et al. 2009), diagenetic
changes of end member isotope values (Macko et al.
1983), and the photosynthetic discrimination relative to
water that results in the depletion of the H isotope
deuterium in phytoplankton.