Ecological Entomology (2007), 32, 45–52
© 2007 The Authors
Journal compilation © 2007 The Royal Entomological Society 45
Introduction
Introductions of exotic plants and phytophagous insects to new
geographic areas can result in the rapid evolution of novel
plant–insect associations (Strong, 1974; Berenbaum & Zangerl,
1991; Singer et al., 1992; Thompson, 1998). When insects are
introduced, either inadvertently or purposely for the biological
control of exotic weeds, our ability to predict which native
plants are at risk of being colonised by the insect is critical.
Relatedness of a novel host to the usual host plant is generally
thought to be the primary predictor driving host range expan-
sion. Classical biological control of weeds typically involves
evaluating a potential control agent’s performance on native
plant species related to the target weed, under the assumption
that plants closely related to the target are the most vulnerable to
a potential host shift (van Klinken & Edwards, 2002). The re-
stricted host associations highlighted by Ehrlich and Raven
(1964) for several groups of phytophagous insects underscores
the importance of phylogenetic conservatism in host-shifting.
However, lack of congruence commonly seen between the phy-
logenies of host-plant clades and those of the insects that feed
on them (Mitter et al., 1991; Dobler et al., 1996; Becerra &
Correspondence: Naomi Cappuccino, Department of Biology, Carleton
University, Ottawa, Ontario, K1S 5B6 Canada. E-mail: ncappucc@ccs.
carleton.ca
Potential novel hosts for the lily leaf beetle Lilioceris
lilii Scopoli (Coleoptera: Chrysomelidae) in eastern
North America
CRYSTAL ERNST
1
, NAOMI CAPPUCCINO
1
and JOHN THOR
ARNASON
2 1
Department of Biology, Carleton University, Ottawa, Ontario, K1S 5B6 Canada,
2
Department of Biology, University of Ottawa, Ottawa, Ontario, K1N 6N5 Canada
Abstract. 1. Introduced insects often incorporate native plants into their diets and
might be expected to show a predilection for novel hosts that are phylogenetically
related to their normal hosts. The lily leaf beetle, Lilioceris lilii (Coleoptera:
Chrysomelidae), is an introduced pest of cultivated lilies. Oviposition behaviour, larval
behaviour, and development of L. lilii was examined on a range of potential host plants,
as well as on the normal host, Asiatic hybrid lilies Lilium sp.
2. Neonate larval feeding behaviour was quantified on 15 food plant species: 10 from
the Liliales, three from the Asparagales and two eudicots. Larvae fed plants closely
related to the genus Lilium were more likely to initiate feeding, less likely to abandon
their food leaf, and consumed more leaf area.
3. In no-choice tests, females oviposited on the novel hosts Lilium philadelphicum,
Medeola virginiana, Clintonia borealis, Streptopus amplexifolius, and Polygonatum
biflorum; however, all but L. philadelphicum received very few eggs. Non- Lilium novel
hosts were not used for oviposition when presented along with Asiatic lilies in choice
tests.
4. A single individual was reared to the adult stage on the novel host S. amplexifolius.
Several larvae survived to the pupal stage on M. virginiana, although no adults emerged
from those pupae. Larvae reared on the native wood lily L. philadelphicum performed
equally well or better than on the Asiatic cultivar.
5. Our results indicate that the lily leaf beetle poses a threat to native Liliaceae.
Several native Lilium species, including L. philadelphicum, are threatened or endangered
in certain jurisdictions throughout their range; these species should be monitored closely
for colonisation by the beetle.
Key words. Exotic insects, host shifts, host range expansion, Liliales.