Journal oflmmunological Methods, 160 (1993) 267-271 267
© 1993 Elsevier Science Publishers B.V. All rights reserved 0022-1759/93/$06.00
JIM06657
Improved bioassay for the detection of porcine tumor necrosis
factor using a homologous cell line: PK(15)
Giuseppe Bertoni, Peter Kuhnert, Ernst Peterhans and Urs Pauli
Institute of Veterinary I, Trology, University of Bern, Liinggass-Str. 122, CH-3012 Bern, Switzerland
(Received 17 August 1992, revised received 16 November 1992, accepted 30 November 1992)
Close similarities of various physiological parameters makes the pig one of the preferred animal models
for the study of human diseases, especially those involving the cardiovascular system. Unfortunately, the
use of pig models to study diseases such as viral hemorrhagic fevers and endotoxic shock syndrome have
been hampered by the lack of the necessary immunological tools to measure important immunoregula-
tory cytokines such as tumor necrosis factor (TNF). Here we describe a TNF-bioassay which is based on
the porcine kidney cell line PK(15). Compared to the widely used murine fibroblastoid cell line L929, the
PK(15) cell line displays a 100-1000-fold higher sensitivity for porcine TNF-a, a higher sensitivity for
human TNF-a, and a slightly lower sensitivity for murine TNF-a. Using a PK(15) bioassay we can detect
recombinant TNF-a as well as cytotoxic activity in the supernatants of lipopolysaccharide (LPS)-activated
porcine monocytes at high dilutions. This suggests that the sensitivity of the test should permit the
detection of TNF in biological specimens such as pig serum.
Key words: Tumor necrosis factor, porcine; Bioassay; Septic shock
Introduction
Tumor necrosis factor (TNF)-a and -13 are
closely related cytokines. They bind to the same
receptors (TNF-R55 and TNF-R75) and trigger
partly identical and partly different biological re-
actions (Aggarwal, 1990; Fiers, 1991). TNF-a and
Correspondence to: U. Pauli, Institute of Veterinary Virol-
ogy, L~inggass-Str. 122, CH-3012 Bern, Switzerland.
Abbreviations: ASF, African swine fever; EMEM, Eagle's
minimal essential medium; FCS, fetal calf serum; LPS,
lipopolysaccharide; MTT, 3-(4,5-dimethylthiazol-2-yl)-2,5-di-
phenyltetrazolium bromide; OD, optical density; PBMC, pe-
ripheral blood mononuclear cells; PBS, phosphate-buffered
saline; TNF, tumor necrosis factor; rhTNF, recombinant hu-
man TNF-a; rmTNF, recombinant murine TNF-a; rpTNF,
recombinant porcine TNF-~.
-/3 play a central role in the host inflammatory
response against viral, bacterial and parasitic in-
fections (Beutler and Cerami, 1989). TNF either
alone or in association with other cytokines such
as IL-1, IL-6 and interferons, have been impli-
cated in the pathogenesis of different diseases
such as septic shock, cerebral malaria and possi-
bly rheumatoid arthritis (Tracey et al., 1987;
M~innel et al., 1987; Cerami and Beutler, 1988;
Grau et al., 1987; Keffer et al., 1991). It has been
suggested that TNF-a could also play a pivotal
role in the pathology of certain viral diseases such
as African swine fever (ASF) and hog cholera
(Peterhans et al., 1988). These are hemorrhagic
diseases of pig and may serve as potential models
for similar disorders in humans.
In order to develop a pig model to investigate
the role played by TNF-o~ in hemorrhagic viral