Pharmacology Biochemistry & Behavior, Vol. 37, pp. 417--424. © Pergamon Press plc, 1990. Printed in the U.S.A. 0091-3057/90 $3.00 + .00
Inhibition by Antibiotic Tetracyclines
of Rat Cortical Noradrenergic
Adenylate Cyclase and
Amphetamine-Induced Hyperactivity
O. KOFMAN, 1 E. KLEIN, M. NEWMAN, R. HAMBURGER,
O. KIMCHI, T. NIR, H. SHIMON AND R. H. BELMAKER
Beer Sheva Mental Health Centre, Ben Gurion University of the Negev, Beer Sheva
and Ezrath Nashim Hospital, P.O.B. 140, Jerusalem, Israel
Received 6 February 1990
KOFMAN, O., E. KLEIN, M. NEWMAN, R. HAMBURGER, O. KIMCHI, T. NIR, H. SHIMON AND R. H. BELMAKER.
Inhibition by antibiotic tetracyclinesof rat cortical noradrenergicadenylate cyclase and amphetamine-induced hyperactivity. PHAR-
MACOL BIOCHEM BEHAV 37(3) 417--424, 1990.--Two antibiotic tetracyclines, demeclocycline (DMC) and minocycline, share
several biochemical and behavioral properties with lithium (Li). DMC inhibited both noradrenaline- and chloradenosine-sensitive
cyclic AMP accumulation in rat cerebral cortical slices both in vitro and ex vivo following two weeks of chronic dietary treatment.
Minocycline, a lipophilic tetracycline, produced similar results in vitro. Both DMC and minocycline reduced open-field activity
levels in rats following acute treatment, four hours prior to testing. Moreover, both drugs inhibited amphetamine-induced hyperac-
tivity in the open field. Chronic treatment with 0.4% and 0.8% dietary DMC for two weeks attenuated amphetamine hyperactivity
without affecting baseline activity levels in the open field. Neither DMC nor minocycline attenuated apomorphine-indueed stereo-
typy at doses that attenuated amphetamine hyperactivity, a profile which is similar to that of lithium. Unlike lithium, however, DMC
did not reverse reserpine-induced hypoactivity.
Lithium Adenylate cyclase Amphetamine Demeclocycline Minocycline Bipolar affective illness
AMONG Li's numerous biological effects, its inhibition of ade-
nylate cyclase has been extensively studied (3). Li at near
therapeutic concentrations inhibits noradrenaline-sensitive adeny-
late cyclase in rat cortical slices (15). Li at more clearly therapeu-
tic concentrations inhibits noradrenaline-sensitive adenylate cyclase
in slices of human cortex from the normal edges of surgically
removed brain tumours (32). Li at therapeutic concentrations, in
patients under chronic treatment, inhibits the epinephrine-induced
rise in plasma cyclic AMP (13).
Recently, the G-protein component of the adenylate cyclase
complex has been suggested as a molecular site of the Li effect on
adenylate cyclase (1). Since these effects on adenylate cyclase are
unlike those of other psychoactive drugs, compounds with similar
properties should be screened to determine if they have the
behavioral and clinical profile of Li.
Antibiotic tetracyclines have been shown to inhibit ADH-
sensitive adenylate cyclase in the kidney at a site distal to the ADH
receptor (12). This inhibition is similar to that involved in the
pathogenesis of the Li side effect of nephrogenic diabetes insipi-
dus (19, 39, 40). Since adenylate cyclase is similar in many tis-
sues, tetracycline effects on rat cortical noradrenergic adenylate
cyclase were examined in the present study. Since Li inhibits hy-
peractivity induced by low dose amphetamine in rats (7,31), we
also tested the effects of tetracyclines on this behavior. Prelimi-
nary data in both the biochemical and behavioral tests have been
previously published (4,5).
Neuroleptics, but not Li, inhibit apomorphine-indnced stereo-
typy (14); therefore, the effects of tetracyclines on apomorphine-
induced stereotypy were tested to define the specificity of their
behavioral effects. Li has been reported to inhibit reserpine-in-
duced hypoactivity (30) and so here too tetracyclines were stud-
ied. Demeclocycline (DMC) was chosen for most experiments,
since it is the tetracycline with most marked effects on polyuria
and adenylate cyclase in the kidney (18); minocycline was used
in some confirmatory experiments since it is lipophilic and crosses
the blood-brain barrier more readily than DMC (38).
IRequests for reprints should be addressed to Dr. O. Kofman, P.O.B. 4600, Beer Sheva, Israel.
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