ApplMicrobiol Biotechnol (1988) 29:536-543
Applied
Microbiology
Biotechnology
© Springer-Verlag 1988
Kinetics of multiproduct acidogenic
and solventogenic batch fermentations
Mustafa Ozilgen
Food EngineeringDepartment, Middle East TechnicalUniversity,06531 Ankara, Turkey
Summary. Large amounts of data indicated that
most of the metabolic processes of the acidogenic
(acid producing) and the solventogenic (solvent
producing) fermentations were regulated by prod-
uct accumulation. A simple unstructured model
simulated microbial growth, product formation
and substrate utilization in six different fermenta-
tions, where five different microorganisms pro-
duced various combinations of ten different prod-
ucts. Specific growth rates of these microorgan-
isms decreased proportionally with overall prod-
uct accumulation. The products were excreted in
non-growth associated pattern. Excretion of some
of these products were inhibited by the overall
product accumulation similarly as the microbial
growth. A substrate consumption model which
considered the biomass and individually all the
products as separate substrate sinks simulated the
data satisfactorily.
Introduction
In a batch fermenter the specific growth rate of
the microorganisms is generally defined as:
1 dX
~-X dt (1)
where/.t is the specific growth rate, X is the micro-
bial concentration and t is time. The parameter/.t
is generally related to the substrate concentration
with the well known Monod equation (Bailey and
Ollis 1977; Wang et al. 1979):
S
1-,£ = ~rnax Ks..]_S (2)
where ~tma x is the maximum specific growth rate,
Ks is a constant and S is the substrate concentra-
tion. Another widely used expression for the pa-
rameter # is the logistic equation (Weiss and Ollis
1980; Klimek and Ollis 1980; Lam and Ollis
1981; Ozilgen 1988):
~t=~s 1 Xmax (3)
where #s and Xmax are the initial specific growth
rate and the maximum attainable biomass con-
centration, respectively. In Equation 3 the term
(1--X/Xmax) represents slow-down of the growth
with over-crowding of the medium. As the value
of the parameter X approaches Xm~x the logistic
equation predicts no growth (i. e. dX/dt = 0). Inhi-
bitory product accumulation and competition for
the limited nutrient are among the reasons of the
slow-down of the growth in the over-crowded fer-
menter.
Acetate, butyrate, lactate and citric acid are
among the most commonly studied acidic micro-
bial products. Butanol, acetone, ethanol, sorbitol
and butanediol are well known microbial sol-
vents. Ethanol accumulation in the fermenters
(and in the cells) was reported to inhibit various
metabolic activities, such as specific growth rate,
specific ethanol production rate, cell viability and
sugar consumption (Novak et al. 1981 ; Ghose and
Tyagi 1979; Lourerio and van Uden 1982; Leao
and van Uden 1982). Ethanol accumulates in the
cells as a consequence of production and kills
them (Novak et al. 1981). Ethanol stress changes
the lipid composition of the plasma membranes
of the yeasts (Beaven et al. 1982; Thomas et al.
1978; Thomas and Rose 1979). It also inhibits
maltose and glucose transport accross the plasma
membrane (Thomas and Rose 1979).