                  !" ! #  $%  &  ’( " )*  +  )  "  ,   - .  / 0 1   $ )2   3 2               !  "# $ %  &’ (  ) *+, &-’ .$.   /0 1   &’ 2% 32&4 5      5%  .1 /6 (7 ’/ 8 3(4  /0 9  /% &/  /%  % %’  *7%: ’  7% .;.  /0 ! </’    )/%      ) 3)4  (7 (7% ! =$ <  5 )0 = ’      % <!!$. # &’0 > ’?    (  2    @ 9=$= /0 .   )/% 7% (7 / (7     ’ 2  )%7 < 32)<4 .;. /  8% /0 A  8 2% B      5 "’ /  8 B C &D E     34    7 3@4    34  ; !  "#0 5 B F9 $ =; 1>10 G B F9 $ =; 1;!! H0%I/0   %’ */%J  %  , 34 :  *7%: ’J  7% 3<4 +70 & @  */ ;;$1 .;. */  8% 34 /0 5B F19 ;1 $= > $0 G B F19 ;1 $= >$ ;0 ’I80 456 <)+ ’ +#’K  ?K  /  +’/   Research Article Synapse DOI 10.1002/syn.21959 This article has been accepted for publication and undergone full peer review but has not been through the copyediting, typesetting, pagination and proofreading process which may lead to differences between this version and the Version of Record. Please cite this article as an ‘Accepted Article’, doi: 10.1002/syn.21959 © 2017 Wiley Periodicals, Inc. Received: Sep 30, 2016; Revised: Jan 10, 2017; Accepted: Jan 11, 2017 This article is protected by copyright. All rights reserved.