937
Accepted by P. Jäger: 31 Mar. 2005; published: 12 Apr. 2005
1
ZOOTAXA
ISSN 1175-5326 (print edition)
ISSN 1175-5334 (online edition)
Copyright © 2005 Magnolia Press
Zootaxa 937: 1–12 (2005)
www.mapress.com/zootaxa/
Four new species and new records for the spider genus Synotaxus
Simon, 1895 (Araneae: Synotaxidae) from Brazil
ADALBERTO J. SANTOS
1
& CRISTINA A. RHEIMS
1,2
1
Laboratório de Artrópodes, Instituto Butantan. Av. Vital Brasil 1500, CEP 05503-900, São Paulo, SP, Brazil;
oxyopes@yahoo.com
2
Pós-graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo, São Paulo, SP, Brazil;
cris.rheims@butantan.gov.br
Abstract
Four new species of Synotaxus Simon are described and illustrated based on material from northern
and northeastern Brazil: S. brescoviti n. sp., S. bonaldoi n. sp., S. siolii n. sp. and S. itabaiana n. sp.
New records are presented for S. longicaudatus (Keyserling), S. monoceros (Caporiacco) and S.
waiwai Agnarsson. A map with all species records included is provided.
Key words: Taxonomy, zoogeography, distribution range, Neotropical region
Introduction
The spider genus Synotaxus was described by Simon (1895) to include two species of
small, Argyrodes-like Neotropical spiders: S. turbinatus Simon and S. uncatus Simon. The
genus was originally placed in Theridiidae (Simon 1895, Levi & Levi 1962, Exline & Levi
1965). Forster et al. (1990), however, suggested that Synotaxus is related to several genera
of small spiders of Austral temperate distribution, including Physoglenes Simon, a genus
with a problematic history of familial placements. This group was hypothesized as sister to
Theridiidae+Nesticidae, and received the name Synotaxidae, based on Simon’s (1895)
tribal-group Synotaxeae. The monophyly of Synotaxidae, as well as its phylogenetic prox-
imity to the Theridiidae-Nesticidae clade was later corroborated by Griswold et al. (1998)
and Agnarsson (2003, 2004).
According to Agnarsson (2003), Synotaxus monophyly is supported by at least 10
morphological and behavioral characters. This genus can be easily recognized, and sepa-
rated from similar araneoids, by its characteristics “chicken-wire” shaped web (Eberhard
1977: figs. 1, 3; Eberhard 1995: fig. 1; Agnarsson 2003: fig. 6, 2004: fig. 95A) and by the