Physiology & Behavior, Vol. 33, pp. 499-501.Copyright© PergamonPress Ltd., 1984.Printed in the U.S.A. 0031-9384/84$3.00 + .00
BRIEF COMMUNICATION
Nociceptive Thresholds Following Food
Restriction and Return to Free-Feeding
ROBERT J. HAMM AND BRUCE G. LYETH
Department of Psychology, Virginia Commonwealth University, Richmond, VA 23284
Received 15 November 1983
HAMM, R. J. AND B. G. LYETH. Nociceptive thresholds following food restriction and return to free-feeding.
PHYSIOL BEHAV 33(3) 499-501, 1984.--Nine rats were placed on a restricted food diet for 14 days and nociceptive
thresholds were measured by the tall-flick procedure. After 24 hr of food restriction nociceptive thresholds increased. This
initial increase in nociceptive threshold was followed by a decrease in pain threshold on the second day of food restriction.
Nociceptive thresholds returned to pre-deprivation levels on the remaining 11 days of food restriction. When the rats were
given free access to food after 14 days of food restriction, nociceptive thresholds increased 24 hr after the reintroduction of
food and decreased during the next 24 hr. Thus, the results of the present experiment demonstrate that both food restriction
and a rgXuI:n to free feeding after 14 days of food restriction produced the same biphasic pattern of changes in nociceptive
thresholds.
Nociceptive threshold Analgesia Pain Food restriction Starvation Free-feeding Stress
STRESSFUL stimuli produce a number of autonomic ad-
justments in organisms including changes in respiration,
body temperature, metabolic rate, and other neuroendocrine
responses. Recent research has demonstrated that another
component of the stress reaction may be a reduction in sen-
sitivity to pain. For example, exposure to a wide variety of
stressors such as inescapable electric shock [1], immobiliza-
tion [2], rotation [7], and cold water [5] have all produced a
reduction in pain sensitivity following stimulation. Acute
starvation has also been shown to produce a change in
nociceptive thresholds [3]. Starvation produced an increase
in nociceptive threshold that was evident for up to 36 hr.
This period of decreased pain sensitivity was followed by a
decrease in nociceptive threshold. After three days of com-
plete food deprivation, rats were given free access to food.
Nociceptive thresholds were not reliably influenced by a re-
turn to free-feeding conditions. Thus, these results demon-
strate that the onset of starvation produces a biphasic change
in pain sensitivity, and feasting after a short period of star-
vation did not change nociception. Changes in dynorphin, an
endogenous opiate peptide implicated in the control of feed-
ing, has also been examined as a function of food and water
deprivation [11]. As was the case in the Bodnar et al. [3]
study, dynorphin levels were sampled only during the first
three days of complete food and water deprivation. They
found that dynorphin concentrations (measured during the
daytime) increased in the hypothalamus as deprivation
proceeded.
The purpose of the present experiment is to investigate
the longer-term consequences of food restriction on no-
ciception than the three days previously used [3,11]. In
addition, after the animals have adapted to and stabilized on
a restricted food diet, free access to food will be given. The
rationale for reintroducing food after a period of prolonged
deprivation comes from research that has shown that or-
ganisms lower their homeostatic set point for metabolic rate,
body temperature, and respiratory quotient in response to
prolonged fasting [5,10]. The introduction of free access to
food when contrasted with a lowered set point for these
physiological parameters may be reflected in a change in
nociceptive threshold. Thus, the return to free-feeding con-
ditions after a long period of food restriction may produce a
change in pain sensitivity not detected after only three days
of food deprivation.
METHOD
Subjects
Subjects were nine male Sprague-Dawley rats which were
75 days old at the start of the experiment. The average free-
feeding weight of the animals was 355 g. Animals were indi-
vidually housed and water was freely available. The animal
room had a 0600-on/2000-off light-dark cycle.
Apparatus
To measure pain threshold sensitivity, an Emdie tail-flick
apparatus (F'r-6, S/N 2035) was used which consisted of a
421/2× 121/2× 10 cm metal box in which a photo-circuit was
located. Mounted on top of the box was an 8 cm metal plate
in which a shallow groove was cut in which the rat's tail
rested. A small (15 mm) light sensitive optical fiber was lo-
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