Pergamon
Pharmacology Biochemistry and Behavior,Vol.48, No. 1, pp. 1=8, 1994
Copyright ©1994Elsevier Science Ltd
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Antagonist-Precipitated Opioid Withdrawal in
Rats: Evidence for Dissociations Between
Physical and Motivational Signs
GUY A. HIGGINS I AND EDWARD M. SELLERS
Addiction Research Foundation and Departments of Pharmacology and Medicine,
University of Toronto, Toronto, Ontario, Canada
Received 21 December 1992
HIGGINS, G. A. AND E. M. SELLERS. Antagonistprecipitated opioid withdrawalin rats: Evidence for dissociations
betweenphysical and motivationalsigns. PHARMACOL BIOCHEM BEHAV 48(1) 1-8, 1994.-In rats made opioid de-
pendant by the implantation of a single morphine 75 mg base pellet, an attempt was made to determine whether any
correlation existed between physical and motivational withdrawal signs by adjusting the dose of naloxone used to precipitate
withdrawal. The models used to study motivational signs were taste (one- and two-bottle) conditioning and operant responding
for food under an FRI5 schedule of reinforcement. Naloxone at doses of 0.01 mg/kg and above produced both a conditioned
taste aversion (two-bottle test only) and reduced food responding in morphine pellet, but not placebo pellet, implanted
animals. No physical withdrawal signs, e.g., wet dog shakes, diarrhoea, were noted until naloxone doses of 0.05 mg/kg and
above were used. It is concluded that the difference in naloxone doses required to elicit physical and motivational withdrawal
components provides further support for their dissociation.
Naloxone-precipitatedopioid withdrawal Taste conditioning
Physical withdrawal signs Motivational withdrawal signs
Operant responding Rat
EARLY views that physical dependence served as the primary
impetus for substance abuse behavior has been largely super-
seded by an acknowledgement that the positive reinforcing
properties of the abused drugs themselves serve a critical role
in this process [e.g., see (20,37,38) for reviews]. Indeed, by
implication, adaptive processes such as physical dependence
are subsequent to chronic drug use and, therefore, offer no
explanation as to why this behavior becomes established in
the first place. Nevertheless, within certain drug classes, par-
ticularly the opiates, the induction of physical dependence and
consequent avoidance of an aversive withdrawal state, repre-
sents one aspect of the motivational impetus maintaining drug
self-administration behavior (18,37). It may be hypothesized
then, that drugs that relieve withdrawal could serve as useful
therapies for the treatment of substance, including opiate,
abusers.
Before attempting to identify such agents it is important to
have some knowledge as to the clinical manifestations of a
drug withdrawal state. Opioid withdrawal is characterized by
a wide range of symptoms including nausea, gastrointestinal
disturbance, various sympathomimetic reactions, agitation/
anxiety, and marked dysphoria (18,19,39). The time course
and severity of these various symptoms are variable between
patients and dependent on a variety of factors such as the
particular drug, dally dosage, duration of use, environmental
context, and method of withdrawal (i.e., spontaneous or an-
tagonist-precipitated withdrawal) (18). However, irrespective
of this, it seems to be a common observation that this phe-
nomenon has profound behavioral, psychological, and physi-
ological components.
In preclinical research there has been a greater emphasis
towards the delineation of these aspects of opioid withdrawal,
for it seems that multiple systems are implicated. The peri-
aqueductal grey and locus coeruleus appear to be particularly
important for the expression of many physical opioid with-
drawal signs (4,24,25) although other brain/spinal areas have
been implicated (23-25) and a peripheral component may be
responsible for secretory signs such as diarrhoea (23,25,27).
s Requests for reprints should be addressed to Guy A. Higgins, Glaxo Unit of Behavioural Psychopharmacology, University of Hertfordshire,
College Lane, Hatfield, Herts, UK.