XPD Polymorphism and Risk of Subsequent Cancer in
Individuals with Nonmelanoma Skin Cancer
Abenaa M. Brewster,
1
Anthony J. Alberg,
2
Paul T. Strickland,
3
Sandy C. Hoffman,
2
and Kathy Helzlsouer
2
1
Department of Clinical Cancer Prevention, University of Texas M.D. Anderson Cancer Center, Houston, Texas and Departments of
2
Epidemiology and
3
Environmental Health Sciences, Johns Hopkins Bloomberg School of Public Health, Johns Hopkins University,
Baltimore, Maryland
Abstract
Background: Individuals with nonmelanoma skin
cancer (NMSC) are at increased risk of developing
subsequent cancers. Genetic predisposition to reduced
DNA repair capacity may be an underlying suscepti-
bility factor explaining the excess risk of malignancies.
To test this hypothesis, a cohort study was conducted
to examine the association between XPD Lys751Gln
polymorphism and risk of a second primary cancer in
individuals with NMSC. Methods: A subgroup of 481
individuals with a history of NMSC who participated
in the CLUE II community-based cohort was followed
for the development of a second primary cancer. Blood
specimens donated in 1989 were genotyped for the XPD
Lys751Gln polymorphism using the 5V nuclease assay.
Cox proportional regression with delayed entry was
used to calculate the incidence rate ratio (IRR) and 95%
confidence interval (95% CI) for risk of developing a
second primary cancer according to XPD genotype. All
statistical tests were two sided. Results: Eighty indi-
viduals developed a second primary cancer. The most
frequent occurring cancers were of the prostate (18%),
lung (15%), and breast (15%). Persons with at least one
Gln allele had an increased risk of a second primary
cancer compared with the reference Lys/Lys genotype
(adjusted IRR 2.22, 95% CI 1.30-3.76). When the
reference category was limited to never smokers with
the Lys/Lys genotype, the risk of developing a second
primary cancer associated with having at least one Gln
allele was increased >3-fold in both never smokers
(IRR 3.93, 95% CI 1.36-11.36) and ever smokers (IRR
6.14, 95% CI 2.17-17.37). Conclusion: These findings
suggest that individuals with NMSC who have the
variant XPD Gln allele are at increased risk of devel-
oping a second primary cancer. (Cancer Epidemiol
Biomarkers Prev 2004;13(8):1271 – 5)
Background
Nonmelanoma skin cancers (NMSC) are the most
common cancers diagnosed in the United States. Over 1
million new cases of basal and squamous cell carcinomas
of the skin occur annually, and exposure to UV radiation
is the leading risk factor (1, 2). UV radiation causes skin
carcinogenesis by inducing cyclobutane pyrimidine
dimers and pyrimidine (6-4) pyrimidone photoproducts
that form between adjacent pyrimidine bases (3).
Suboptimal repair of these DNA lesions by the nucleo-
tide excision repair (NER) pathway leads to the
accumulation of mutations in important cell cycle
regulatory genes that may eventually result in tumor
development (4, 5).
The overall prognosis of NMSC is excellent, but
several studies have shown that affected individuals
are at increased risk of developing subsequent cutaneous
and noncutaneous malignancies (6-19) and have a higher
overall cancer mortality rate (20, 21) compared with the
general population. The excess incidence of noncuta-
neous malignancies in individuals with a prior NMSC is
unexplained. One hypothesis is that genetic predisposi-
tion to reduced DNA repair capacity may be an
underlying susceptibility factor, increasing the risk of a
subsequent malignancy (19, 22).
Polymorphisms of genes involved in the major NER
pathway are of particular interest as candidate genes that
may contribute to interindividual variations in DNA
repair capacity and cancer susceptibility. In addition to
the removal of mutagenic UV-induced photoproducts,
the NER pathway is responsible for the removal of bulky
and helical distorting DNA adducts induced by chemical
carcinogens and cellular metabolites (3, 23).
Given the involvement of the NER pathway in the
removal and repair of a wide variety of carcinogenic
DNA lesions, defects in this pathway may predispose an
individual to both cutaneous and noncutaneous malig-
nancies. Within this pathway, a plausible candidate sus-
ceptibility gene is the common XPD polymorphism in
exon 23 that causes an amino acid substitution of lysine
for glutamine (24). The normal functioning XPD protein
Received 12/29/03; revised 3/3/04; accepted 3/11/04.
Grant support: National Institute of Aging grant 5U01AG018033, National Cancer
Institute grant 5U01CA086308, and National Institute of Environmental Health
Sciences grant P30 ES03819. Minority Supplement of Clinical Oncology Research
Career Development Program (K12-CA01709) from the National Cancer Institute
(A.M. Brewster) and KO7 award (CA73790) from the National Cancer Institute
(A.J. Alberg).
The costs of publication of this article were defrayed in part by the payment of
page charges. This article must therefore be hereby marked advertisement
in accordance with 18 U.S.C. Section 1734 solely to indicate this fact.
Requests for reprints: Kathy Helzlsouer, Department of Epidemiology, Johns
Hopkins Bloomberg School of Public Health, Johns Hopkins University, Suite E6132,
615 North Wolfe Street, Baltimore, MD 21205. Phone: 410-955-9727; Fax: 410-614-2632.
E-mail: khelzlso@jhsph.edu
Copyright D 2004 American Association for Cancer Research.
Cancer Epidemiology, Biomarkers & Prevention 1271
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