Int. J. Vitam. Nutr. Res. 80 (4 – 5), 2010, 307 – 313 307
DOI 10.1024/0300-9831/a000038 Int. J. Vitam. Nutr. Res. 80 (4 – 5) © 2010 Hans Huber Publishers, Hogrefe AG, Bern
Article to the Special Issue
The Use of Caco-2 Cells to
Estimate Fe Absorption in
Humans – a Critical Appraisal
Ann-Sofie Sandberg
Chalmers University of Technology, Department of Chemical and Biological Engineering, Food Science, Gothenburg, Sweden
Abstract: The Caco-2 cell model is widely used to assess the bioaccessibility/availability of iron from
foods and diets. Analysis of iron uptake in this human epithelial cell line is usually preceded by a two-
step digestion to simulate the conditions in the stomach and small intestine. Moreover, culturing the cells
on inserts permits the measurement of iron transport. The cellular iron uptake is determined by direct
measurements using radioisotopes, or indirectly by measurement of ferritin, the intracellular storage
form of iron. There is a good correlation between Caco-2 cell uptake and human iron bioavailability
for a number of dietary factors known to affect iron absorption. However, recent data suggest that in
some cases there is no correlation. Possible reasons for such discrepancies, the benefits, and limitations
of the Caco-2 cell model are discussed. In conclusion, in vitro experiments with Caco-2 cells are impor-
tant tools for ranking foods with respect to bioavailability, for mechanistic studies of iron absorption,
and for studies of dietary factors influencing absorption. The results need to be confirmed in humans.
Key words: Caco-2 cells, iron bioavailability, in vitro digestion
Introduction
Low bioavailability of iron is a major nutritional prob-
lem in developing countries, where the diet is based on
staple foods. One strategy to improve the iron status
is to biofortify staple food crops with iron and other
micronutrients. For this reason, to rank the foods with
respect to iron bioavailability, there is a need for a sim-
ple method to estimate iron bioavailability in humans.
Such a method must be reproducible and reflect the
in vivo conditions. It is thus of great importance that
the in vitro methods are validated by in vivo measure-
ments in humans. Traditionally a combined two step
in vitro digestion and measurement of dialyzable or
soluble iron is used to determine iron availability [1];
i. e. , iron that is potentially available for absorption in
the gut. Dialyzability methods are useful for screening
purposes to identify major differences in availability,
but it cannot predict the magnitude of response [2, 3].
For example, methods based on dialyzability exclude
iron that is bound to large but not small molecules.
However, some of these iron-bound large molecule
complexes (e. g., ferritin) have substantial bioavail-
ability whereas iron bound to certain small molecules
is not bioavailable [4].
Combining in vitro digestion with uptake in Caco-2
cells is a step forward since it predicts both availabil-
ity and uptake into the enterocyte and at times also
the absorption. However, in a consensus statement of
experts working in the field of iron nutrition, it was