EDITORIAL COMMENTARY
The atrial neural network: Ablation minefield or strategic
target?
Kevin F. Kwaku, MD, PhD, Richard L. Verrier, PhD
From the Cardiovascular Division, Beth Israel Deaconess Medical Center, and Harvard Medical School, Boston, MA.
Background
Neural control of heart rhythm is achieved through complex
hierarchal inputs from both the extrinsic autonomic nervous
system and the less well understood intrinsic cardiac neural
network.
1,2
The latter system consists of an extensive set of
neuron types, including afferent sensory fibers as well as
sympathetic and parasympathetic efferents. The intrinsic
neural network exerts local control of chronotropy, dromot-
ropy, and inotropy mediated by ganglionated plexi (GP) in
discrete epicardial fat pads initially characterized by the
pioneering work of Randall,
3
Zipes,
4
and Armour,
1,2
and
their respective coworkers. Several major fat pads have
been located in proximity to the atria in mammalian species.
The sinoatrial (SA) node fat pad is located at the caudal end
of the SA node and contains many of the anterior right (AR)
GP. The atrioventricular (AV) node fat pad is located at the
junction of the inferior vena cava with both atria and con-
tains ganglia from the inferior right (IR) GP. The superior
left pulmonary vein (PV) fat pad is proximate to that PV and
contains the superior left (SL) GP, as depicted in Fig. 1 of
Hou et al.
5
In addition, there is yet another distinct fat pad
in canines, which is located between the medial superior
vena cava (SVC) and aortic root in proximity to the right
pulmonary artery.
4
The human heart also contains epicardial fat pads
6
and an
extensive distribution of GPs
1
. The location of these struc-
tures generally corresponds to that observed in canines in
their proximity to the atria and major vessels.
1,2,6
Cum-
mings and colleagues
7
have recently reported on the rele-
vance of an anterior fat pad, located between the base of the
aorta (Ao) and pulmonary artery in humans, which may
correspond to the canine SVC-Ao fat pad. Thus, while fat
pad locations may not be identical, there appear to be
general interspecies homologies in cardiac neural architec-
ture.
Functional studies indicate that disruption of the GPs has
the potential for enhancing electrical inhomogeneity, lead-
ing to arrhythmias.
4
In a recent clinical study, it was dem-
onstrated that preservation of the anterior fat pad signifi-
cantly decreases the incidence of postoperative atrial
fibrillation (AF).
7
Conversely, selective ablative targeting of
GPs has been advanced by some centers as a useful adjunct
to pulmonary vein isolation in the treatment of AF.
8,9
These
incongruous observations underscore the importance of ad-
vancing our understanding of the GP network.
Main findings and commentary
The present study by Hou and coworkers
5
advances an
emerging theme that the GPs, which exhibit extensive an-
atomical interconnections, function as an interactive net-
work in their influence on the SA and AV nodes. The
specific hypothesis tested was that as a result of connectivity
within the atrial neural network, GPs remote from the SA
and AV nodes modulate the effects of GPs more proximate
to these structures. The investigators studied the AR, IR,
and SL GPs in anesthetized, open-chest canines.
A systematic combination of electrical stimulation of the
GPs was performed with a handheld bipolar electrical probe
in conjunction with blockade of adjoining GPs either with
lidocaine or radiofrequency ablation. They measured the
responses in sinus and ventricular rates and the atrial-His
intervals. From their findings, the investigators deduced that
IR GP control of AV nodal dromotropy is modulated by
remote inputs from both the AR and SL GP. Similarly, the
AR GP control of sinus rate is modulated by remote inputs
from the SL GP but not the IR GP. In addition, a weaker,
direct influence of the SL GP on the SA node was observed.
Overall, the authors concluded that AR GP and IR GP serve
integrative roles in modulating SA and AV node function,
as depicted in their Fig. 5.
5
The principle emanating from these observations is that
the atrial neural network operates through an interactive
hierarchy among the GPs with respect to relative influence
on the SA and AV nodes. While this basic premise appears
sound, a number of caveats need to be stated. The model
explores the interactions among three important GPs, but
the authors acknowledge that investigation of potential par-
ticipation of the SVC-Ao fat pad is not incorporated in their
model. This is an important limitation, since this structure
appears to receive the majority of efferent vagal innervation
Address reprint requests and correspondence: Richard L. Verrier,
Ph.D., Associate Professor of Medicine, Harvard Medical School, Beth
Israel Deaconess Medical Center, Harvard-Thorndike Electrophysiology
Institute, Harvard Institutes of Medicine, 77 Avenue Louis Pasteur, Room
223, Boston MA 02115. E-mail : rverrier@bidmc.harvard.edu.
1547-5271/$ -see front matter © 2007 Heart Rhythm Society. All rights reserved. doi:10.1016/j.hrthm.2006.10.006