LEAD ARTICLE
Distinct Xp11.2 Breakpoint Regions in Synovial Sarcoma
Revealed by Metaphase and Interphase FISH:
Relationship to Histologic Subtypes
B. de Leeuw, R. F. Suijkerbuijk, D. Olde Weghuis, A. M. Meloni,
G. Stenman, L. G. Kindblom, M. Balemans, E. van den Berg,
W. M. Molenaar, A. A. Sandberg, and A. Geurts van Kessel
ABSTRACt: F/uorescence in situ hybridization (FISH) and molecular analyses ofsynovial sarcomas with
cytogenetically similar (X;18)(p11.2;q11.2) translocations have revealed two alternative breakpoint regions
in Xp11.2, one residing in the ornithine aminotrunsferase-like I (OATL1) region and the other one in the
related but distinct OATL2 region. As these results were obtained by different groups, we set out to evalu-
ate an extended series of tumors with special emphasis on the two possible X-related breakpoint regions.
Together, seven synovial sarcomas were identified with a break in the OATL1 region and six with a break
near OATL2, thereby confirming the actual existence of the two alternative Xp breakpoint regions. We
speculate that there seems to be a relationship between the occurrence of these breakpoint regions and
the histologic phenotype of the tumors, with a predominance ofOATLl-related breakpoints in the classi-
ca• biphasic tumors and of OATL2-related breakpoints in the monophasic fibrous tumors.
INTRODUCTION
Synovial sarcoma is a soft tissue tumor which occurs mainly
in adolescents and young adults. Next to rhabdomyosarcoma,
synovial sarcoma is the most common solid tumor in chil-
dren under 18 years of age [1, 2]. The patients' ages range
from 11 months to 78 years, whereas the mean age at the time
of operation is 37 years [3, 4]. About 90% of the primary
tumors are found in the extremities, especially the lower
limbs [2], whereas 3-10% are found in the head and neck
region, heart, and mediastinum [5-7]. In spite of its name,
which arose from the light microscopic resemblance of the
classic biphasic type to synovial tissue, the exact histogenetic
origin of synovial sarcoma is, as yet, unknown. Electron mi-
croscopic and immunologic data clearly distinguish the cells
from true synovial cells [8, 9]. Also, the tumors may arise
at places where normally synovial cells are not present [10].
From the Department of Human Genetics (13. d L., R. F. S.,
D. O. W.., M. B., A. G. v K.), University Hospital, Nijmegen, The
Netherlands; Southwest Biomedical Research Institute of Genetr/x
Inc. (,4. M. M., A. A. S.), Scottsdale Arizona; Laboratory of Cyto-
genetics (G. S., L. G. K.), Department of Oral Pathology, University
of G~teborg, Sweden; Department of Medical Genetics (E. v d B.),
University of Groningen, The Netherlands; and Department of
Pathology (W. M. M.), University of Groningen, The Netherlands.
Address repr/nt requests to: B. de Leeuw, Department of Human
Genetics, University Hospital, P.O. Box 9101, 6500 HB Nijmegen,
The Netherlands.
Received July 12, 1993; accepted September 3, 1993.
For this reason suggestions have been made to change the
name to carcinosarcoma [8, 9].
Histologically, synovial sarcomas can be divided into two
major groups: the classical biphasic type, containing an epi-
thelial and a spindle cell component, and the monophasic
type, containing only spindle cells, although some areas may
express epithelial markers like EMA, keratin, CEA, $100, and
others [8, 9]. The tumors differ in a graded fashion, how-
ever, which has led some authors to subdivide the biphasic
type into totally biphasic (or glandlike) and focally biphasic
[11]. Occasionally, two other types of synovial sarcoma are
mentioned, namely the undifferentiated type and the mono-
phasic epithelial type. These latter tumors are presumed to
represent extreme and rare cases of monophasic and biphasic
synovial sarcomas, respectively [1, 11]. The monophasic spin-
dle cell and biphasic subtypes are thought to occur with simi-
lar frequencies [12], although some authors state that the
monophasic type is predominant, comprising about 60% of
all cases [9].
A characteristic chromosomal translocation, t(X;18)(p11.2;
q11.2], has been found in synovial sarcoma cells [13]. Some-
times this translocation is the only cytogenetic abnormality
present, which is indicative of a primary causal event in tu-
mor development. Recently, we [14, 15] and others [16] have
identified yeast artificial chromosomes CYACs) containing hu-
man genomic fragments that span the breakpoint region on
the X chromosome in different tumors. Both YACs (number
2 and 7) contain ornithine amino transferase (OAT)dike se-
quences (OATL1 and OATL2, respectively] that are located
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