Ugolini et al. 1 Chemical Extraction of Microorganisms From 1 Water-Saturated, Packed Sediment 2 3 Fabio Ugolini 1 , Martin H. Schroth 1 *, Helmut Bürgmann 2 , Frederik Hammes 3 , Josef Zeyer 1 4 5 6 1 Institute for Biogeochemistry and Pollutant Dynamics, ETH Zürich, Universitätstrasse 16, 7 CH- 8092 Zürich, Switzerland 8 2 Eawag, Swiss Federal Institute for Aquatic Science and Technology, Department of Surface 9 Waters - Research and Management, CH-6047 Kastanienbaum, Switzerland 10 3 Eawag, Swiss Federal Institute for Aquatic Science and Technology, Department of 11 Environmental Microbiology, CH-8600 Dübendorf, Switzerland 12 *Institute for Biogeochemistry and Pollutant Dynamics, ETH Zürich, Universitätstrasse 16, 13 CH- 8092 Zürich, Switzerland. e-mail: martin.schroth@env.ethz.ch. Corresponding author. 14 15 16 Research paper submitted to 17 Water Environment Research 18 May 31, 2012 19 Revised October 12, 2012 20 21