Introduction
Relaxin (RLX) is a peptide hormone secreted mainly by the
corpus luteum during pregnancy [1], with well established
effects on the female reproductive organs. Cardiomyocytes
from rat atria have been shown to secrete detectable amounts
of RLX suggesting the cardiovascular system as a physiolog-
ical target [2]. In fact, RLX has a powerful vasodilatatory
action on rat uterus and mesocaecum, mouse mammary
gland and pigeon crop sac [3], and decreases blood pressure
in spontaneous hypertensive rats [4]. In some vascular
districts, such as the coronary arteries of the guinea pig, the
RLX induced increase in coronary flow is accounted for by
the generation of nitric oxide (NO) [5].
The NO donors, sodium nitroprusside and glyceryl tri-
nitrate, down regulate the immunological and non-immuno-
logical release of histamine from sensitised tissues of the
guinea pig [6]. RLX shares, with the NO donors the ability
to down regulate the immunological and non-immunological
release of histamine from isolated rat and guinea pig mast
cells [7]. Cardiac anaphylaxis in vitro is a reliable model of
immune responses [8] in which the changes in myocardial
functions are attributed to the release of pro-inflammatory
and vasoactive mediators from cardiac mast cells. Therefore,
we were interested to extend previous investigations [7] con-
cerning the interactions between RLX and the immune
response to the isolated heart. Our present aim was to cor-
relate the effects of RLX on cardiac anaphylaxis with the
generation of NO in the isolated heart of the guinea pig.
Materials and methods
Guinea pig hearts, taken 18–25 days after sensitisation by intraperito-
neal and subcutaneous injections of egg albumin (1 ml of 1 % solution),
were perfused with Tyrode’s solution at 37°C in a Langendorff ap-
paratus and challenged with egg albumin (0.1 ml of 1 % solution into the
aortic cannula). Records were taken of the strength of contraction, rate
and coronary outflow. RLX was added to the perfusion fluid 60 min
before antigen challenge. Histamine released in the perfusates and re-
Inflamm. res. 50, Supplement 2 (2001) S 122 – S 123
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© Birkhäuser Verlag, Basel, 2001
Inflammation Research
Relaxin generates nitric oxide and provides protection against
cardiac anaphylaxis
J. F. Ndisang
1
, R. Baronti
1
, G. Cecere
1
, E. Masini
1
, D. Bani
2
and P. F. Mannaioni
1
1
Department of Preclinical and Clinical Pharmacology, Viale G. Pieraccini n° 6, 50139 Florence, Italy, Fax : +39 055 427 1280,
e-mail: mannaion@pharm.unifi.it
2
Department of Anatomy, Histology and Forensic Medicine, University of Florence, Viale G. Pieraccini 6, 50139 Florence, Italy
Correspondence to: P. F. Mannaioni
tained in the heart was detected fluorimetrically. In cardiac homo-
genates, cyclic GMP levels were detected by radioimmunoassay using
125
I-labelled cyclic GMP. Tissue Ca
2+
levels were determined by means
of atomic absorption.
Results and discussion
We have previously shown that RLX increases both the
coronary flow in the isolated heart of the guinea pig and the
amount of nitrite that appears in the perfusates. Both effects
were significantly reduced by pretreatment with the nitric
oxide synthase inhibitor N-monomethyl-L-arginine, and
were comparable with those obtained by the endothelium-
dependent vasodilator acetylcholine as well as by the endo-
thelium independent vasodilator, sodium nitroprusside [5].
The present experiments show that in the presence of
RLX the pathophysiological responses to antigen challenge
were fully abated (Fig. 1, panel a). Consistent eg with these
observations, the release of histamine induced by antigen
was also diminished in the presence of RLX (Fig. 1, panel b).
Moreover, RLX significantly increased cGMP levels and
reduced the antigen-induced increase in cardiac calcium
(data not shown). It should be noted that in the presence of
RLX, antigen challenge evoked a less sustained coronary
constriction and a more pronounced coronary dilatation (data
not shown).
The results of these experiments show that RLX, at a
concentration as low as that measured in plasma under
physiological conditions, provides protection against cardiac
anaphylaxis, in that the responses to antigen are fully abated
and the release of histamine substantially reduced. These
effects are concomitant with the increase in cardiac cGMP
and with the corresponding decline in the immunological rise
of cardiac calcium. It is conceivable that the protection af-
forded by RLX on cardiac anaphylaxis could be accounted
for by the generation of NO. In fact, the immunological and
non-immunological release of histamine from isolated mast
cells is similarly reduced by RLX in a NO-dependent
fashion. The inhibition of mast cell histamine release by RLX
is consistent with the formation of NO, which actually occurs
from rat mast cells in the presence of RLX [7], as well as