1 23 Zoomorphology Evolutionary, Comparative and Functional Morphology ISSN 0720-213X Zoomorphology DOI 10.1007/s00435-013-0200-9 Comparative anatomy of the heart– glomerulus complex of Cephalodiscus gracilis (Pterobranchia): structure, function, and phylogenetic implications Sebastian Merker, Alexander Gruhl & Thomas Stach