Acute Effects of Microcystis aeruginosa from the Patos Lagoon Estuary, Southern
Brazil, on the Microcrustacean Kalliapseudes schubartii (Crustacea: Tanaidacea)
W. Montagnolli, A. Zamboni, R. Luvizotto-Santos, J. S. Yunes
Unidade de Pesquisas em Cianobacte ´rias (UPC), Departamento de Quimica, Fundac ¸a ˜o Universidade Federal do Rio Grande (FURG), Cx Postal 474,
CEP 96201-900, Rio Grande/RS, Brazil
Received: 12 January 2003 /Accepted: 6 September 2003
Abstract. Toxic blooms of the cyanobacterium Microcystis
aeruginosa, a microcystin producer, have been observed in the
past two decades in the Patos Lagoon estuary (southern Brazil).
This cyanobacterium reaches the estuary from northern waters
and accumulates as toxic blooms in the shallow margins of the
environment. Microcystins are phosphatase (PP1 and PP2A)
inhibitors and cause animal death via alteration of the liver cell
cytoskeletons and intrahepatic hemorrhage. The massive accu-
mulation of toxic material affects the survival of several ben-
thonic estuarine local organisms. The tanaidacea Kalliapseudes
schubartii is a benthonic estuarine species which occurs at high
densities throughout the year in mixohaline areas of the Patos
Lagoon. This microcrustacean is of high ecological relevance
and plays an important role in the estuarine food web, as it is
consumed on a large scale by estuarine fish. This work verifies
the acute toxicity of aqueous extracts of M. aeruginosa
RST9501 and of sediments spiked with lyophilized material of
the same strain on K. schubartii; it also evaluates the sublethal
effects on tanaidacean oxygen consumption rates and glycogen
levels under acute exposure to M. aeruginosa aqueous extracts.
The strain M. aeruginosa RST9501 was cultured in BGN/2
medium. The aqueous extracts were prepared using the lyo-
philized material from the strain cultures. Acute tests were
performed over 96 h at a salinity of 15, at six toxic concentra-
tions, and resulted in an average 96-h LC50 of 1.44 mg ml
-1
.
The spiked sediment tests were performed with a 10-day du-
ration, using the lyophilized material in three proportions of
powder/sediment and showed an average LC50 of 1.79 mg
ml
-1
. Oxygen consumption was determined after 24 and 48 h
of incubation in adult organisms exposed to sublethal aqueous
extract concentrations and showed a significant increase at the
highest concentrations. This suggests alterations in the organ-
ism’s metabolism by exposure to the cyanobacterium extract.
The glycogen levels were determined with a commercial kit
(Glicox 500; DOLES Ltd.); after 24 and 48 h the dosages were
administered in the same organisms utilized in the oxygen
consumption test and did not demonstrate significant differ-
ences. The results demonstrate the possible risks of intoxica-
tion to which the natural populations of K. schubartii were
exposed in the environment and emphasize the importance of
studies involving sublethal concentrations of M. aeruginosa to
other organisms of the trophic web in this aquatic system.
Cyanobacterial blooms have several consequences for water
quality and their collapse frequently causes high mortality
among aquatic animal populations (Vasconcelos et al. 2001).
Among many species of cyanobacteria that can develop toxic
blooms, Microcystis aeruginosa is one of the most common
and is a matter of great concern (Chorus and Bartram 1999).
The cyanobacterium Microcystis aeruginosa produces toxins
called microcystins, monocyclic heptapeptide molecules, of
which over 65 structural variants are known. Most of the
variants are potent hepatotoxins and tumor promoters in mam-
mals (Codd et al. 1999; Sivonen and Jones 1999). At the
molecular level, microcystins are serine/threonine phosphatase
PP1 and PP2A inhibitors, resulting in hyperphosphorylation of
proteins, affecting intracellular signaling, cell growth, and dif-
ferentiation processes, and inducing cell disturbance (Toivola
and Eriksson 1999).
Previous works have shown that microcystin can be conju-
gated with the tripeptide glutathione (GSH), through a conju-
gation reaction catalyzed by glutathione-S-transferase (GST)
(Pflugmacher et al. 1998; Takenaka 2001; Vinagre et al. 2003).
The microcystin–GSH complex has a less inhibitory effect on
phosphatases, clearly showing that this conjugation reaction is
involved in detoxification processes (Metcalf et al. 2000). The
enzyme (GST) is reported to be involved in detoxification
reactions of several kinds of pollutants, such as metals, fungi-
cides, and pesticides (Freedman et al. 1989; Gallagher et al.
1992; Maracine and Segner 1998).
Different aquatic organism species from all trophic levels in
the web are susceptible to microcystins, these include bacteria,
protozoa, macrophytes, green algae and diatoms, and micro-
and mesozooplankton, as well as invertebrates and fish (Christ-
offersen 1996). A number of reports have demonstrated the
sensitivity of different species of marine and freshwater crus-
tacea to Microcystis aeruginosa cells and microcystins
(Delaney and Wilkins 1995; Salomon et al. 1996; DeMott Correspondence to: J. S. Yunes; email: dqmsarks@furg.br
Arch. Environ. Contam. Toxicol. 46, 463– 469 (2004)
DOI: 10.1007/s00244-003-2304-6
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