Physiology& Behavior, Vol. 47, pp. 601-603. ©Pergamon Press plc, 1990. Printed in the U.S.A. 0031-9384/90 $3.00 + .00
BRIEF COMMUNICATION
Effects of Preferential Delta and
Kappa Opioid Receptor Agonists on the
Intake of Hypotonic Saline
BLAKE A. GOSNELL, MARK J. MAJCHRZAK AND DEAN D. KRAHN
University of Michigan, Department of Psychiatry, University Hospital, 8D8806, Box 0116
Ann Arbor, MI 48109-0116
Received 16 November 1989
GOSNELL, B. A., M. J. MAJCHRZAK AND D. D. KRAHN. Effects of preferential delta and kappa opioid receptor agonists on
the intake ofhypotonic saline. PHYSIOL BEHAV 47(3) 601-603, 1990. - - A previous study has implicated central mu opioid receptors
in the preference for salt solutions. Because mu, kappa and delta receptors are all thought to play a role in food intake and/or the
mediation of palatability, we performed a series of experiments to determine whether preferential agonists at kappa and delta receptors
might also stimulate the intake of salt solutions. When injected centrally into nondeprived rats, two selective agonists at delta receptors
caused increases in the intake of 0.6% saline; the intake of concurrently available water was either unchanged or slightly increased.
The selective kappa agonist U-50,488H had no effect on water or saline intake, whereas the preferential kappa agonist DAFPHEDYN
caused a delayed increase in saline intake. These results indicate a role for central delta receptors in the preference for salt solutions,
and are consistent with the suggestion that opioids play a role in the mediation of palatability.
Opioids Palatability Taste preference Reward Drinking Saline intake
SEVERAL reports have suggested that endogenous opioids play a
role in mediating palatibility [see (5,12) for reviews]. Generally,
opioid antagonists have been found to decrease the preference for
(or intake of) palatable fluids (3, 5, 8), while opioid agonists
increase preferences (2,8). Although most studies have dealt with
the consumption or preferences for sweet-tasting fluids, there are
indications that these opioid effects can also be obtained with
nonsweet palatable fluids. Hypo- and isotonic salt solutions are
preferred over water even in salt-replete rats (10), which suggests
that these solutions are palatable to rats. Naloxone caused a
selective reduction in the intake of hypotonic saline solutions (4),
and reduced 0.9% saline intake at lower doses than those required
to reduce water intake (11). In water-deprived rats, systemic
injections of morphine increased rats' preference for isotonic
saline (1).
There are at least three types of opioid receptors, the best
characterized being the mu, kappa and delta (6). Recently, we
found that a selective mu agonist stimulates the intake of hypo-
tonic (0.6%) saline in nondeprived rats (9). Because mu, kappa
and delta receptors are all thought to play a role in food intake
and/or the mediation of palatability (5, 7-9), we performed a
series of experiments to determine whether preferential agonists at
kappa and delta receptors might also stimulate the intake of salt
solutions. The delta agonists used were [D-Thr 2, LeuS]enkephalin -
Thr 6 (DTLET) and [D-Pen 2, D-PenS]enkephalin (DPDPE);
the kappa agonists were U-50,488H and a recently re-
ported dynorphin analog, [D-Ala2, (Fs)Phea]-dynorphinl_13-NH2
(DAFPHEDYN) (13).
METHOD
Male Sprague-Dawley rats (Charles River Laboratories, Inc.,
Wilmington, MA) were used in all experiments. At the time of
surgery, body weights ranged from 239-345 g. They were
individually housed in stainless steel cages, and food and water
were available ad lib, except as described below. Lights were on
for 12 hours daily, and all experiments were carried out in the light
period. A 20-gauge cannula was implanted into the fight lateral
ventricle of each rat with techniques similar to those previously
reported (8,9). Cannula placement was verified by the observation
of a drinking response to an ICV injection of angiotensin II (AII,
100 ng). All rats were tested at least once with AII, and all data
from animals drinking less than 5 ml in 30 min on all tests were
excluded. Adaptation to testing procedures began no sooner than 3
days after surgery.
After an overnight period of water deprivation, rats were
moved to individual test cages for adaptation to the testing
procedure. One hour later, a 0.6% sodium chloride (saline)
solution was presented to each rat in a 25-ml graduated buret fitted
with a stainless steel sipper tube. After two hr of access to this
solution, rats were returned to their home cages. After this first
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