Developmental Brain Research, 38 (1988) 161-166 161
Elsevier
BRD 50669
Research Reports
Effects of short-term prenatal alcohol exposure on
neuronal membrane order in rats
Charles V. Vorhees 1, Scott Rauch 2, and Robert Hitzemann 1
llnstitute for Developmental Research, Children's Hospital Research Foundation, and Department of Pediatrics, University of
Cincinnati, Cincinnati, 0H45229 (U.S.A.) and 2Department of Psychiatry, University of Cincinnati, Cincinnati, 0H45267 (U.S.A.)
(Accepted 7 July 1987)
Key words: Prenatal alcohol; Brain neuronal membrane order; Brain membrane fluidity and development;
Ethanol and brain neuronal membrane
Long-Evans rat dams were treated with ethanol (4 g/kg, twice daily) by gavage on gestational days 10-14. This dosage schedule has
been shown to produce significant behavioral and ponderal teratogenicity. Pair-fed dams were gavaged with isocaloric amounts of su-
crose. All offspring were reared by untreated, surrogate dams. Pups were sacrificed on days 3 and 28, and whole brain neuronal plas-
ma membranes were prepared for analysis by a fluorescence polarization technique using 1,6-diphenyl-l,3,5-hexatriene as the mem-
brane probe. On day 3, steady-state anisotropy was significantly decreased in the ethanol-treated pups. Arrhenius plots revealed that
this difference was associated with a change on both membrane entropy and enthalpy. By day 28, the differences between groups dis-
appeared. These data would be consistent with the view that the brief gestational ethanol exposure delays neuronal maturation.
INTRODUCTION
It is now clear that alcohol is both a behavior-
al 1'2'29'37'40 and structural teratogen H,25,32. The be-
havioral effects can be seen at alcohol exposure lev-
els below those required for overt structural malfor-
mations. Some of the most common behavioral ef-
fects of fetal alcohol exposure have included in-
creased activity or reduced locomotor habituation
using the open-field 6,7,9A°,17,18,30 and holeboard34; re-
duced avoidance performance using conditioned
taste aversion 3-s, one and two-way active avoid-
ance 8'38, and passive avoidance3,16,35; increased reac-
tivity using auditory startle4; reduced appetitive dis-
criminationS; increased perseverative behavior using
T-maze learning or spontaneous alternation36; and
delayed reflex and physical development using tests
of n¢gative geotaxis, surface righting, and eye open-
ing2~,28, 38.
Recently, we have examined the behavioral and
ponderal teratogenicity associated with a discrete al-
cohol exposure during days 10-14 of gestation 41.
Ethanol-exposed offspring exhibited delayed olfac-
tory orientation to home cage scent and delayed low-
er incisor eruption compared to pair-fed or ad lib fed
controls. After weaning, the ethanol offspring exhi-
bited increased open-field activity, particularly of
centrally located sections, and facilitated swimming
performance in a water maze. Ethanol-exposure sig-
nificantly decreased weight gain and increased post-
natal, but not prenatal, mortality in the progeny. The
female ethanol offspring also showed delayed vaginal
patency development.
The biochemical mechanisms underlying the be-
havioral abnormalities associated with short and/or
long-term prenatal ethanol-exposure are unknown.
Current theories suggest that ethanol acts by disrupt-
ing (perturbing) the neuronal membrane lipid ma-
Correspondence: R. Hitzemann. Current address: Department of Psychiatry and Behavioral Science, State University of New York
at Stony Brook, Health Sciences Center, Stony Brook, NY 11794-8101, U.S.A.
0165-3806/88/$03.50© 1988 Elsevier Science Publishers B.V. (Biomedical Division)