Vol 62 International Journal of Dairy Technology 1
ORIGINAL
RESEARCH
*Author for
correspondence. E-mail:
desmond.morgan@ucd.ie
© 2008 Society of
Dairy Technology
doi: 10.1111/j.1471-0307.2008.00435.x
Blackwell Publishing Ltd Oxford, UK IDT International Journal of Dairy Technology 1364-727X 1364-0307 Society of Dairy Technology 2008 XXX ORIGINAL RESEARCH ORIGINAL RESEARCH
Effect of high intensity pulsed electric fields on enzymes
and vitamins in bovine raw milk
JOERG RIENER, FRANCESCO NOCI, DENIS A CRONIN,
DESMOND J MORGAN* and JAMES G LYNG
UCD School of Agriculture Food Science and Veterinary Medicine, College of Life Sciences, UCD Dublin, Belfield,
Dublin 4, Ireland
Pulsed electric fields (PEF) were applied to fresh bovine raw milk using a laboratory-scale continuous
PEF system to study the impact on selected native enzyme activities and on vitamin degradation. The
activities of protease, lipase, alkaline phosphatase and lactoperoxidase following PEF treatment at
electric field strengths ranging from 15 to 35 kV/cm for treatment times of 12.5 μs to 75 μs were
examined. Reductions in enzyme activity, even under the most severe conditions employed, were quite
modest, amounting to 14%, 37% and 29% for lipase, protease and alkaline phosphatase, respectively,
while the activity of lactoperoxidase remained unchanged. The PEF treatment had no effect on the levels
of thiamin, riboflavin, retinol and α-tocopherol in the milk.
Keywords Bovine milk, Enzymes, Pulsed electric fields, Vitamins.
*Author for correspondence. E-mail: desmond.morgan@ucd.ie
INTRODUCTION
Generally, fresh unpasteurized raw milk is heat
treated to guarantee a microbiologically safe and
shelf-stable product. While thermal treatments
are very efficient, some undesirable organoleptic
and chemical changes can occur. The latter may
include the formation of lactulose, furosine or
hydroxymethylfurfural, as well as some protein
degradation (Morales et al. 1996; Contarini et al.
1997; Lopez-Fandino and Olano 1999; Villamiel
et al. 1999; Calligaris et al. 2004). An interesting
alternative to heat treatment is the application
of pulsed electric fields (PEF), because of its
effectiveness against most microorganisms, while
flavour, colour and vitamins are only minimally
or nonaffected (Sizer and Balasubramaniam 1999;
Cserhalmi et al. 2003).
The effect of PEF on enzyme inactivation has
not been widely investigated and there is a lack
of published information. In milk there are around
70 indigenous enzymes and several of these are
significant in relation to the quality of milk and
milk products (Kelly and Fox 2006). For the present
study, four enzymes that have important roles in
milk quality were selected. Lipases are enzymes
that catalyse the hydrolysis of triglycerides, the
major lipid components of milk. The significance
of lipolysis in milk is twofold: flavour production
and altered functionality (Ha and Lindsay 1993).
Free fatty acids, particularly those of short and
medium chain length, have strong flavours that are
generally considered as undesirable (Balcao and
Malcata 1998; Deeth 2006). The application of 20
pulses at 21.5 kV/cm to lipase in milk led to a complete
inactivation (Grahl and Märkl 1996) (Table 1).
Bendicho et al. (2002b) observed a reduction of
62.1% maximum activity when it was treated at
27.4 kV/cm and 80 pulses. However, when PEF
treatments were applied in a continuous flow
mode by the same research group, an inactivation
rate of just 13% was achieved, after applying 80
pulses at 37.3 kV/cm and 3.5 Hz. Proteases are
enzymes that degrade proteins. Raw milk may have
proteolytic activity of endogenous or indigenous
origin, which can adversely affect milk flavour
(Chen et al. 2003). It was found that PEF reduced
protease activity by 62.7% after an 896 μs treatment
at 35.5 kV/cm and 111 Hz (Bendicho et al. 2003b).
An 80% reduction in protease activity was obtained
in tryptic soy broth with yeast extract after 20
pulses of 2 μs and 18 kV/cm at 0.25 Hz. When the
protease was treated in sterilized skim milk, PEFs
of 14 and 15 kV/cm and pulsing rates of 1 and 2 Hz
achieved 40% and 60% inactivation after 32 and
98 pulses of 2 μs duration, respectively (Vega-
Mercado et al. 1995; Vega-Mercado et al. 1997).
Alkaline phosphatase (ALP) is a native enzyme
found in raw milk. It is a rather nonspecific mono-
phosphoester hydrolase that catalyses the removal
of phosphate groups. Conventionally, the effective-
ness of milk pasteurization processes is monitored
by the inactivation of ALP, which is more heat
resistant than the pathogens present in milk (Mussa