Novel Vancomycin Dimers with Activity against
Vancomycin-Resistant Enterococci
Uma N. Sundram and John H. Griffin*
Department of Chemistry, Stanford UniVersity
Stanford, California 94305-5080
Thalia I. Nicas
Infectious Disease Research
Lilly Research Laboratories, Lilly Corporate Center
Indianapolis, Indiana 46285
ReceiVed June 24, 1996
Vancomycin typifies the glycopeptide family of antibacterial
agents, which exhibit an unusual, receptor-like mode of action:
by binding with high affinity and specificity to the C-terminal
L-lysyl-D-alanyl-D-alanine portion of peptidoglycan precursors,
vancomycin prevents their incorporation into the polymeric
bacterial cell wall.
1
Williams et al. have detailed a second
molecular recognition function of glycopeptides, that of self-
association into homodimers. They have demonstrated that
glycopeptide dimerization can be highly favorable and coopera-
tive with the binding of peptide ligands.
2
They have also
provided evidence that self-association can lead to enhancements
in in Vitro antibacterial potency,
3
consistent with a model in
which dimerization increases the intrinsic affinity of glycopep-
tides for their peptidoglycan precursor targets, and increases
the overall avidity of interaction between self-associated, ditopic
glycopeptide receptors and polyvalent ligands presented by the
bacterial cell.
3,4
Interestingly, vancomycin self-associates in
solution only weakly (K
dim
) 700 M
-1
), suggesting that
dimerization may not play a significant role in the biological
action of this clinically important agent. This prompted us to
examine the effects that coValent dimerization would have on
the molecular recognition and antibacterial properties of van-
comycin. We now report the synthesis of a series of novel bis-
(vancomycin)carboxamides and the discovery that some of these
compounds exhibit promising in Vitro activity against vanco-
mycin-resistant enterococci and selectively enhanced affinity
for depsipeptide ligands mimicking peptidoglycan precursors
from these organisms.
Bis(vancomycin)carboxamides 1a-d (Figure 1) were pre-
pared by HBTU-mediated coupling of vancomycin (2.2 equiv)
with 1,6-diaminohexane, cystamine, homocystamine, and tri-
ethylenetetramine, respectively.
5
The dimers were isolated in
44-68% yield by reversed-phase HPLC and characterized by
high-field
1
H NMR spectroscopy and liquid secondary ion (LSI)
or electrospray (ES) mass spectrometry. As controls, the
monomeric adducts of vancomycin with cystamine (2b) and
triethylenetetramine (2d) were prepared by use of excess amine.
The in Vitro antibacterial properties of vancomycin, 1a-d,
2b, and 2d were determined by broth microdilution assays. It
was found that both monomeric and dimeric vancomycin
carboxamide derivatives retained activity against vancomycin-
susceptible Gram-positive organisms, including staphylococci
and enterococci (Table 1). Strikingly, dimers 1a-c display
substantially enhanced in Vitro potency against strains of
enterococci which exhibit high-leVel resistance to Vancomycin
and other glycopeptides (Vancomycin-resistant enterococci
(VRE, VanA phenotype).
6
The average minimum inhibitory
concentration (MIC) displayed by bis(vancomycin)cystamide
(1b) against VRE (19 μg/mL, 11 μM in vancomycin subunits)
is more than a factor of 60 lower than that displayed by
vancomycin against these strains and approximately 10-fold
higher than the MICs displayed by vancomycin against sus-
ceptible enterococci. Dimerization appears to be required for
significant activity against VRE, since 2b and 2d did not inhibit
(1) (a) Perkins, H. R.; Nieto, M. Pure Appl. Chem. 1973, 35, 371-381.
(b) Reynolds, P. E. Eur. J. Clin. Microbiol. Infect. Dis. 1989, 8, 789-803.
(2) (a) Mackay, J. P.; Gerhard, U.; Beauregard, D. A.; Maplestone, R.
A.; Williams, D. H. J. Am. Chem. Soc. 1994, 116, 4573-4580. (b) Gerhard,
U.; Mackay, J. P.; Maplestone, R. A.; Williams, D. H. J. Am. Chem. Soc.
1993, 115, 232-237. (c) Waltho, J. P.; Williams, D. H. J. Am. Chem. Soc.
1989, 111, 2475-2480.
(3) Beauregard, D. A.; Williams, D. H.; Gwynn, M. N.; Knowles, D. J.
C. Antimicrob. Agents Chemother. 1995, 39, 781-785.
(4) Mackay, J. P.; Gerhard, U.; Beauregard, D. A.; Westwell, M. S.;
Searle, M. S.; Williams, D. H. J. Am. Chem. Soc. 1994, 116, 4581-4590.
(5) Sundram, U. N.; Griffin, J. H. J. Org. Chem. 1995, 60, 1102-1103.
(6) (a) Leclerq, R. E.; Derlot, R. E.; Duval, J.; Courvalin, P. N. Engl. J.
Med. 1988, 319, 157-161. (b) Uttley, A. H. C.; Collins, C. H.; Naidoo, J.;
George, R. C. Lancet 1988, 1, 57-58. (c) Courvalin, P. Antimicrob. Agents
Chemother. 1990, 34, 2291-2296.
Figure 1. Bis(vancomycin)carboxamides 1a-d.
Table 1. In Vitro Antibacterial Properties and Ligand-Binding
Affinities of Monomeric and Dimeric Vancomycin Carboxamide
Derivatives
MIC (μM)
a
Kd
b
(μM)
cmpd
S. aureus
(mean)
c
Enterococcus
(mean)
d
VRE
(mean)
e
Ac2KDADLac Ac2KDADA
Van 0.30 1.0 690 1800 ( 65 1.1 ( 0.4
1a 2.0 0.51 32 180 ( 20 1.5 ( 0.1
1b 0.48 0.50 11 440 ( 70 1.9 ( 0.8
1c 3.2 0.29 15 680 ( 160 3.9 ( 0.6
1d 0.46 2.9 >69 1160 ( 380 2.9 ( 0.4
2b 0.08 1.4 >67 2600 ( 120 4.3 ( 0.1
2d 0.40 2.8 >60 3630 ( 230 8.4 ( 0.9
a
Minimum concentration of derivative required to inhibit growth
of bacterial cells in cation-adjusted Mueller-Hinton broth. For 1a-d,
MIC values refer to the concentrations of individual vancomycin
subunits.
b
Dissociation constants determined by competitive titration
with AcK(F)DADA in 10 mM HEPES, 6 mM NaCl, pH 7.0, buffer.
For 1a-d, Kd values refer to the noncooperative association of
individual vancomycin subunits to the ligands.
c
Average MIC values
for 10 vancomycin-susceptible strains of Staphylococcus aureus.
d
Average MIC values for 4 vancomycin-susceptible strains of Entero-
coccus faecium and Enterococcus faecalis.
e
Average MIC values for
4 strains of E. faecium and E. faecalis exhibiting high-level resistance
to vancomycin (VanA genotype confirmed by PCR).
13107 J. Am. Chem. Soc. 1996, 118, 13107-13108
S0002-7863(96)02129-4 CCC: $12.00 © 1996 American Chemical Society