Research Article
The Effect of Breed, Gender, and Acid Stimulation in
Dog Saliva Proteome
Sónia Lucena,
1,2
Ana V. Coelho ,
3
Fernando Capela-Silva ,
1,4
Asta Tvarijonaviciute ,
5
and Elsa Lamy
1
1
Instituto de Ciˆ encias Agr´ arias e Ambientais Mediterrˆ anicas (ICAAM), Universidade de
´
Evora, 7000-083
´
Evora, Portugal
2
Departamento de Medicina Veterin´ aria, Escola de Ciˆ encias e Tecnologia, Universidade de
´
Evora, 7000-083
´
Evora, Portugal
3
Instituto de Tecnologia Qu´ ımica e Biol´ ogica Ant´ onio Xavier, Universidade Nova de Lisboa, 2780-157 Oeiras, Portugal
4
Departamento de Biologia, Escola de Ciˆ encias e Tecnologia, Universidade de
´
Evora, 7000-671
´
Evora, Portugal
5
Interdisciplinary Laboratory of Clinical Analysis (Interlab-UMU), Regional Campus of International Excellence “Campus Mare
Nostrum”, University of Murcia, 30100 Espinardo, Murcia, Spain
Correspondence should be addressed to Elsa Lamy; ecsl@uevora.pt
Received 10 February 2018; Accepted 4 April 2018; Published 3 June 2018
Academic Editor: Giuseppe Piccione
Copyright © 2018 S´ onia Lucena et al. Tis is an open access article distributed under the Creative Commons Attribution License,
which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly cited.
Saliva gained interest as a potential noninvasive source of biomarkers in humans and that interest starts to be extended also to other
animal species. For this purpose, the knowledge of the salivary proteome in healthy conditions and the factors that afect it and
how they afect it are necessary. Te aim of the present study was to assess the efect that gender and breed have in saliva proteome
and the changes in it induced by stimulation with acid. Saliva from 4 diferent purebred dogs (Portuguese Podengo, Greyhound,
Rafeiro Alentejano, and Beagle) of both genders was collected without and afer stimulation with lemon juice. SDS-PAGE and two-
dimensional gel electrophoresis (2-DE) profles were compared and the proteins of interest in-gel digested and identifed by mass
spectrometry. Acid stimulation decreased total protein concentration and the relative amounts of some protein bands/spots. Gender
appeared to have minimal efect in saliva proteome, whereas the infuence of breed varies. Beagles and Portuguese Podengos were
the two breeds with higher diferences. In conclusion, stimulation procedures and dog breed should be considered in data analysis
when using salivary proteins for diagnostic purposes.
1. Introduction
Physiological variables are of added value to assess the welfare
and lifespan both in humans and in animals, as they pro-
vide important information for interpreting and validating
emotional and biological responses, respectively [1]. Saliva
has gained interest for biomarker identifcation, mainly due
to the noninvasive nature of its collection; at the same time
that it contains glandular and blood-born molecules that
can change under diferent conditions [2]. In dogs, most of
the studies have been focused on the evaluation of stress
by measuring salivary cortisol levels [3]. Infectious agents,
such as Helicobacter spp., Bartonella spp., or rabies virus,
have also been evaluated [4–6]. In addition, canine saliva has
been used for quantifcation of acute phase proteins [7] and
allergen measurements [8] and in forensic studies for canine
mRNA determination [9]. Furthermore, recently, healthy
dog saliva proteome has been characterized by shotgun
proteomics, with the identifcation of 2,491 proteins and
peptides [10]. Despite this characterization, two-dimensional
electrophoresis (2-DE) salivary protein profles of dog saliva
have been less explored. Although several researchers con-
sider that gel-based approaches provide limited information,
2-DE continues providing reliable quantitative results on
diferential protein expressions as they display a high num-
ber of protein species, their isoforms, and posttranslational
modifcations at the same time [11]. It also has the advantage
of allowing modifcations of the protein mixtures caused by
inadequate treatment or endogenous protease activities with
physiological relevance to be easily recognized via pattern
disturbances by 2D gels [11].
Hindawi
BioMed Research International
Volume 2018, Article ID 7456894, 12 pages
https://doi.org/10.1155/2018/7456894