Null Results in Brief
The Role of Established Breast Cancer Susceptibility
Loci in Mammographic Density in Young Women
Eunjung Lee,
1
Christopher A. Haiman,
1
Huiyan Ma,
2
David Van Den Berg,
1
Leslie Bernstein,
1,3
and Giske Ursin
1,4
1
Department of Preventive Medicine, Keck School of Medicine, University of Southern California, USC/Norris Comprehensive
Cancer Center, Los Angeles, California;
2
Institute for Health Promotion and Disease Prevention Research, Keck School of
Medicine, University of Southern California, Alhambra, California;
3
City of Hope Comprehensive Cancer Center and
Beckman Research Institute, Duarte, California; and
4
Department of Nutrition, University of Oslo, Oslo, Norway
Introduction
Recently, three genome-wide association studies identi-
fied and validated multiple novel loci as contributors to
breast cancer susceptibility (1-3). In two of these studies
(1, 2), the most significant association was with a
common variant (rs2981582) in FGFR2 , a gene implicat-
ed in mammary carcinogenesis (4-6). However, for all
loci, the specific biological pathways that are disrupted
and the mechanisms through which these common
genetic variations contribute to breast cancer risk
remain unclear.
Mammographic density (MD) is an important risk
factor for breast cancer (7) and might be predominantly
inherited (8), but the gene(s) responsible are, to a large
extent, unknown. In the present study, we investigated
whether these established breast cancer variants are also
associated with MD. Given the known effects of postme-
nopausal hormone therapy on MD (9, 10), we addressed
this question in a study of breast cancer patients under
theageof50.
Materials and Methods
This study has previously been described (11). Female
patients diagnosed with histologically confirmed first
primary invasive breast cancer were identified through
the Los Angeles County Cancer Surveillance Program.
EligiblecaseswereU.S.-bornandEnglishspeaking,white
(including Hispanic) or African-American females, ages
20to49years.Amongthe2,882potentiallyeligiblecases,
1,794 (62%) were interviewed. The study was approved
by the Institutional Review Board of the University of
Southern California. All participants provided written
informedconsent.
All subjects were interviewed in-person using a
structured questionnaire which covered information on
standard breast cancer risk factors. We obtained and
digitized mammographic films of the contralateral (non-
cancerous) breast on 639 of 866 women with unilateral
cancer for whom we requested mammograms. Estrogen
and progesterone receptor information (ER/PR) was
abstracted from pathology reports.
Bloodspecimenswerecollectedfrom588(92%)women
with mammograms. DNA was available on 578 women
forthecurrentstudy.Theheterogeneityofgeneticeffects
has been observed for some of these variants between
African-Americans and other populations, which pre-
sumably reflects differences in linkage disequilibrium
between these variants and the underlying causal alleles
(1, 3). Thus, we limited our analysis to 516 whites (429
non-Hispanic, 87 Hispanic). We genotyped six single
nucleotide polymorphisms (SNP; rs889312, rs2981582,
rs3803662, rs3817198, rs13281615, and rs13387042) using
TaqMan assays as previously described (1, 3). The
genotyping call rate was 97% to 98% for all six SNPs.
We included 33 blind duplicate samples which had
completely consistent results with the original samples.
All SNPs were in Hardy-Weinberg equilibrium in each
population.
MD was quantified (by G. Ursin) using the University
of Southern California Madena computer-assisted assess-
ment method (12). The breast area was outlined by a
research assistant trained by G. Ursin. The Madena soft-
warecountstheareaofabsolutedensityaswellasthetotal
breast area. The percent mammographic density was
equivalent to the amount of absolute density divided by
thetotalbreastarea.
Statistical Analyses. We examined the association
between the carrier status of these SNPs and the
mammographic percentage density using multivariable
linear regression. The models were adjusted for age at
diagnosis,ethnicity,menopause,hormoneuse,andbody
mass index 1 year prior to the diagnosis. We examined
allele dosage effects because the previous studies
Cancer Epidemiol Biomarkers Prev 2008;17(1). January 2008
Cancer Epidemiol Biomarkers Prev 2008;17(1):258–60
Received 11/2/07; accepted 11/2/07.
Grant support: CA17054 and CA74847 from the National Cancer Institute, NIH,
4PB-0092 from the California Breast Cancer Research Program of the University of
California, and in part through NIH contract no. N01-PC-35139. The collection of
cancer incidence data used in this publication was supported by the California
Department of Health Services as part of the statewide cancer reporting program
mandated by California Health and Safety Code Section 103885. The ideas and
opinions expressed herein are those of the authors, and no endorsement by the State
of California, Department of Health Services is intended or should be inferred.
The costs of publication of this article were defrayed in part by the payment of page
charges. This article must therefore be hereby marked advertisement in accordance
with 18 U.S.C. Section 1734 solely to indicate this fact.
Requests for reprints: Giske Ursin, Department of Preventive Medicine, Keck School
ofMedicine,UniversityofSouthernCalifornia/NorrisComprehensiveCancerCenter,
Room 4407, 1441 Eastlake Avenue, Los Angeles, CA 90089. Phone: 323-865-0423;
Fax: 323-865-0142. E-mail: gursin@usc.edu
Copyright D 2008 American Association for Cancer Research.
doi:10.1158/1055-9965.EPI-07-2749
258
Research.
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