CSIRO PUBLISHING
Volume 49, 1998
© CSIRO Australia 1998
A journal for the publication of original contributions
in physical oceanography, marine chemistry,
marine and estuarine biology and limnology
www.publish.csiro.au/journals/mfr
All enquiries and manuscripts should be directed to
Marine and Freshwater Research
CSIRO PUBLISHING
PO Box 1139 (150 Oxford St)
Collingwood Telephone: 61 3 9662 7618
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Published by CSIRO PUBLISHING
for CSIRO Australia and
the Australian Academy of Science
&
Marine
Freshwater
Research