Journal of Tropical Forest Science 26(4): 532–542 (2014) Dong TL et al.
532 © Forest Research Institute Malaysia
SITE CONDITIONS FOR REGENERATION OF HOPEA ODORATA
IN NATURAL EVERGREEN DIPTEROCARP FOREST IN
SOUTHERN VIETNAM
TL Dong
1, 2
, CL Beadle
1, 3
, R Doyle
1
& D Worledge
3
1
Tasmanian Institute of Agriculture and School of Land and Food, The University of Tasmania, Private Bag 98, Hobart,
Tasmania 7001, Australia
2
Silviculture Research Institute, Vietnamese Academy of Forest Science, Duc Thang, Bac Tu Liem, Hanoi, Vietnam; dong.
tran@vafs.gov.vn
3
CSIRO Ecosystem Sciences, Private Bag 12, Hobart, Tasmania 7001, Australia
Received July 2013
DONG TL, BEADLE CL, DOYLE R & WORLEDGE D. Site conditions for regeneration of Hopea odorata in
natural evergreen dipterocarp forest in southern Vietnam. Matching species to suitable sites is important in
reforestation. This study investigated the site conditions that support regeneration of Hopea odorata, a valuable
timber species, in a secondary evergreen natural forest. Stand structure, light intensity at seedling level and
soil condition were examined in three representative 50 × 50 m plots. The upper canopy was dominated
by four dipterocarps: H. odorata, Shorea roxburghii, Anisoptera costata and Dipterocarpus alatus. The prevailing
stand structure supported vigorous germination but not development of seedlings of all four species. Low
light levels near the forest floor were the major constraint on seedling development of H. odorata. There was
no regeneration when the mean per cent transmitted incident daily photosynthetic active radiation (PAR)
was 2.2%; seedling germination but not development was supported when PAR was 6.6%; regeneration
and development occurred when PAR was 11.4%. The soils were slightly acidic with low clay and high sand
contents and low nutrient concentration, but this was apparently not a constraint on growth given adequate
light conditions. The results suggest that the re-establishment of H. odorata on degraded sites using nurse
crops should be possible provided that high levels of shading are avoided.
Keywords: Gap regeneration, photosynthetically active radiation, site requirement
INTRODUCTION
Valuable tropical timber species have a history
of overharvesting (Lamb 2011). One of these
species is Hopea odorata , a late successional
tropical dipterocarp which is naturally distributed
in many South-East and South Asian countries
(Prosea Foundation 1993). Its timber is valued
because of its durability and resistance to insects,
and use for weight-bearing construction. Hopea
odorata is currently assessed as vulnerable in the
IUCN Red List of threatened species (IUCN
2012). In Vietnam, it is listed as a priority species
in need of immediate conservation (Hong 2012).
Under natural conditions, usually in riparian
and moist forest, H. odorata grows with other
dipterocarps which eventually occupy the
upper storey and become dominant. Natural
regeneration occurs where there is shade,
suggesting that seedlings are shade tolerant
(Sakai et al. 2009). Young trees become more
light demanding (Thao 1995). While shade
is considered important for germination and
initial establishment within the understorey
(Kettle 2009), it is associated with low rates of
growth, rates increasing when light intensities are
increased (Appanah 1998). Continuous shade
can compromise survival (Kamaluddin & Grace
1993).
There have been many attempts using
enrichment planting or planting with nurse
crops to re-establish H. odorata for either timber
or conservation (Weinland 1998). In Vietnam,
it has been a major species in reforestation
programmes, but many of these have failed
because of the use of inappropriate planting sites
and silvicultural treatments (Tam 2007). Nutrient
deficiency and soil compaction led to poor
seedling establishment on degraded rainforest
soils in Sabah (Nussbaum et al. 1995). In lowland