Journal of Water Resource and Protection, 2013, 5, 1117-1126
Published Online November 2013 (http://www.scirp.org/journal/jwarp)
http://dx.doi.org/10.4236/jwarp.2013.511117
Open Access JWARP
Presence of Multidrug-Resistant E. coli, Enterococcus
spp. and Salmonella spp. in Lakes and
Fountains of Porto, Portugal
Carlos E. Flores
1,2*
, Luís Loureiro
3
, Lucinda J. Bessa
4
, Paulo Martins da Costa
1,4
1
ICBAS-Abel Salazar Biomedical Sciences Institute, University of Porto, Porto, Portugal
2
Faculty of Sciences (FCUP), University of Porto, Porto, Portugal
3
Tulane University, School of Public Health and Tropical Medicine, New Orleans, USA
4
CIIMAR-Interdisciplinary Center for Marine and Environmental Research, University of Porto, Porto, Portugal
Email:
*
carloseduardof1990@gmail.com
Received September 8, 2013; revised October 11, 2013; accepted November 6, 2013
Copyright © 2013 Carlos E. Flores et al. This is an open access article distributed under the Creative Commons Attribution License,
which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly cited.
ABSTRACT
Urban lakes and fountains provide recreational activities that could facilitate the contact between humans, animals and
biological agents. The objective of this work was to assess the water quality and safety of 17 lakes and 13 fountains in
the city of Porto (Portugal), by detecting the presence of Escherichia coli, enterococci and Salmonella spp., and analyz-
ing their antimicrobial resistance. The mean number of E. coli colony forming units (CFU) was higher, both in lakes
(2.67 × 10
3
CFU/100 ml) and fountains (3.52 × 10
2
CFU/100 ml), in comparison to enterococci counts: 5.60 × 10
2
and
4.10 × 10 CFU/100 ml, respectively. By screening a set of 26 multidrug-resistant E. coli (including seven ESBL-pro-
ducing isolates) for virulence genes, we found strains carrying the fimH, chuA, cvaC, cnf1, hlyA, TSPE4.C2 and yjaA
genes. Phylogenetic analyses have shown that the majority of E. coli strains belong to groups A and B1. Salmonella spp.
was found in two lake water samples. Generally, the fecal contamination was notorious and worrisome, requiring pre-
ventive measures to minimize the negative impacts on human health. Finding the contamination source is definitely im-
portant so that interventions can be more effective.
Keywords: Antimicrobial Resistance; Enterococcus spp.; Escherichia coli; Salmonella spp.; Fountains; Lakes
1. Introduction
Recreational and decorative use of water can have im-
portant effects on health and well-being. Presently, in se-
veral cities, lakes and fountains are mainly used for these
purposes. However, the contamination of urban lakes and
fountains with fecal material can turn these water spots
into potential hazards for public health. The presence of
total coliforms, fecal coliforms and Escherichia coli is
indicators of fecal pollution and commonly used to ana-
lyze the safety and quality of water supplies and recrea-
tional water [1,2].
E. coli belongs to the family of Enterobacteriaceae
and is a common inhabitant of the gastrointestinal tract of
humans and animals, and can be easily spread through
water contaminated with fecal material. As a genetically
diverse group, most strains of E. coli are harmless com-
mensals, but others are capable of causing either intesti-
nal or extra-intestinal (ExPEC isolates) disease, due to the
existence of virulence genes in their genome or plasmids
[3]. According to Clermont et al. [4], the analysis of vi-
rulence genes contributes to the classification of E. coli
into phylogenetic groups. The most virulent strains be-
long to the phylogenetic group B2, with a minority of
strains belonging to group D. Most of the commensal
strains are included in group A and B1 [5].
Enterococci are also present in the normal gut of ani-
mals and humans, and have been used as a water quality
indicator in many countries. Presently, the genus Ente-
rococcus comprises 51 validly published species [6]. How-
ever, Enterococcus faecium and Enterococcus fecalis are
the most prevalent species in animal feces and commonly
used as indicators of fecal pollution [7,8].
In general, due to the difficulty associated with the di-
rect isolation of pathogens from water samples, indica-
tors of fecal pollution may denounce the presence of pa-
thogenic microorganisms (such as salmonella) in water
*
Corresponding author.