Trabecular anisotropy and collagen fibre orien-
tation in the mandibular condyle following
experimental functional appliance treatment
using sheep
B. Ma
1,6
, W. J. Sampson
1,6
, O. W. Wiebkin
2,6
, D. F. Wilson
3,6
, N. L. Fazzalari
4–6
¹Orthodontic Unit, Dental School;
2
Department of Medicine, Royal Adelaide Hospital;
3
Oral Pathology, Dental School;
5
Department of Pathology;
6
Australian Jaw Joint Project,
Faculty of Health Sciences, The University of Adelaide, Adelaide, Australia
4
Division of Tissue Pathology, Institute of Medical and Veterinary Science, Adelaide, Australia
Summary
In order to study the modifying effects of functional ap-
pliances on the mechanical environment of the tempo-
romandibular joint (TMJ), we characterised the struc-
ture of the mandibular condyle subsequent to an ex-
perimental functional appliance intervention. Eight,
four-month-old, castrated male Merino sheep, were
randomly allocated to experimental and control groups
(n = 4 in each group). Forward mandibular displace-
ment was induced with an intraoral appliance. The
study period was 15 weeks, during which time fluor-
ochromes were administered to all of the animals. Mid-
sagittal sections of the TMJ were selected for analysis
and trabecular anisotropy was estimated using bone
histomorphometry. Only the experimental group dem-
onstrated that the trabecular bone in the central condy-
lar region was less anisotropic when compared to the
subchondral region. Also, the variation in trabecular
anisotropy of the central condylar region was found to
be smaller in the experimental group. The collagen
fibre orientation was analysed under polarised light as
the proportion of the dark or bright fibres observed in
regions which existed before, and regions which formed
during the experiment, as determined by the fluoroch-
rome labels. In the experimental group, more bright
collagen fibres were found in the most superior region
of the mandibular condyle when compared with the
controls. These results suggested that the experimental
functional appliances changed the orientation and pat-
tern of the mechanical forces acting on the mandibular
condyle, and possibly increased the magnitude of the
lateral functional forces applied to the most superior
part of the condyle during such treatments.
Keywords
Sheep mandibular condyle, growth modification,
collagen fibre orientation, trabecular anisotropy
Vet Comp Orthop Traumatol 2006; 19: 35–42
Received November 20, 2004
Accepted July 20, 2005 Vet Comp Orthop Traumatol 1/2006
Original Research 35
© 2006 Schattauer GmbH
Introduction
Growth modification in the region of the
human temporomandibular joint (TMJ) by
the use of functional appliances is of great
interest to orthodontists for the treatment of
class II malocclusion. Functional ap-
pliances have been reported to achieve cor-
rection of class II discrepancy through in-
creasing the mandibular length (17, 29, 34)
and rotating the mandible (6, 35). These
changes have been suggested to result both
from stimulation of mandibular condylar
growth beyond that which would normally
occur in growing children, and redirection
of condylar growth from an upward and for-
ward vector to a more posterior orientation
(36). However, the mandibular trabecular
bone structural orientation and the ortho-
paedic mechanical stress acting on bone
formation in the TMJ has not been deter-
mined.
The mandibular condyle is a growth site
within the TMJ but current knowledge of
the responses of the mandibular condylar
tissue to functional appliances comes
mainly from animal experiments. In order to
mimic orthodontic treatment procedures,
specific functional appliances have been
used to prompt the mandible to a protrusive
position in various animal species (9, 22,
26). The overall results of the subsequent
bone adaptation have shown increased
bone-forming activity of the mandibular
condylar tissue. For example: this increased
bone-forming activity has been manifested
as elevated alkaline phosphatase activity
and
45
Ca uptake (30) and as newly-formed
trabecular bone being deposited on the pos-
terior border of the condyle and mandibular
ramus (26, 32). In addition, the increased
bone forming activity is not uniformly dis-
tributed among regions within the mandibu-
lar condyle (23).
The bone structure is an indicator of its
mechanical properties having adapted to its
mechanical environment. In trabecular
bone, the anisotropy and bone volume (BV)
to total tissue volume (TV) fraction (BV/
TV) are of special interest, since a large pro-
portion of the variance in the mechanical
properties can be explained by these par-
ameters (24). However, previous reports on
the trabecular architecture of the mandibu-
lar condyle have not provided sufficient in-
formation on how trabecular bone structural
orientation adapts to orthopaedic mech-
anical stress altered by a functional ap-
pliance. After experimental functional ap-
pliance treatment, a posteriorly directed ro-
tation of the trabeculae in the mandibular
condyle has been found (9). More recently,
the BV/TV has been found to be lower in the
mandibular condyle after the treatment des-
pite an increased bone forming activity (23).
Based on available data, functional ap-
pliance treatment seems to change the
orientation of the functional forces acting
on the TMJ whereas the change of the mag-
nitude of the functional force is still uncer-
tain. The same uncertainty regarding
changes in the muscle forces following
functional appliance treatment has also
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