- transition from invasive species control to native species promotion establishment - 131
Applied Vegetation Science 11: 131-138, 2008
doi: 10.3170/2007-7-18335, published online 7 December 2007
© IAVS; Opulus Press Uppsala.
Abstract
Question: Does the seed density of invasive species affect
establishment by native species in a bare ground context
(following invasive species control efforts), and is it possible
to promote transition to a native species dominated state by
manipulating sowing density of the native community?
Location: Experimental wetland basin in Chanhassen, Min-
nesota, USA.
Methods: A mesocosm experiment investigated the infuence
of Phalaris arundinacea (invasive species) propagule pressure
on establishment of native wet meadow species in the context
of a newly restored wetland. Mesocosms were sown with P.
arundinacea (0, 10, 50, 100, or 500 seeds/m
2
) and a mix of
native species (3000 or 15000 seeds/m
2
).
Results: When planted at densities > 100 seeds/m
2
, P. arun-
dinacea increased suppression of native species. Also, high
native seed density suppressed P. arundinacea biomass produc-
tion. This effect was more pronounced when P. arundinacea
seed density was high (> 100 seeds/m
2
), but high native seed
density (15000 seeds/m
2
) did not suppress recruitment of P.
arundinacea from seed.
Conclusions: The transition from post-control bare ground
(a common result of efforts to control invasive species) to
native species establishment depends on both native spe-
cies and invader seed density. These results suggest that a
threshold of P. arundinacea propagule pressure exists, beyond
which transition to a native community is less likely without
management intervention. P. arundinacea can establish in the
presence of a newly developing native plant community, even
at very low densities of P. arundinacea seed. Invader control
(following initial site clearing efforts) is essential to native
species establishment.
Keywords: Colonization; Phalaris arundinacea; Propagule
pressure; Restoration; Revegetation; Sedge meadow; Seed
bank; Threshold dynamics; Wetland.
Nomenclature: Gleason & Cronquist (1991).
The transition from invasive species control to native species promotion
and its dependence on seed density thresholds
Reinhardt Adams, Carrie
*
& Galatowitsch, Susan M.
University of Minnesota, Department of Horticultural Science, 305 Alderman Hall, 1970 Folwell Avenue, St. Paul, MN
55108, USA;
*
Corresponding author; present address University of Florida, Environmental Horticulture Department, PO
Box 110675, Gainesville, FL 32611-0675, USA; Tel. +1 352392183; Fax +1 3523921413; E-mail creinhardt@ifas.uf.edu;
Introduction
Dominance by invasive species prevents the estab-
lishment of target native communities and is a common
barrier to achieving restoration objectives (Kusler &
Kentula 1990; McKinstry & Anderson 1994; Galatow-
itsch et al. 1999a). Thus, invasive species control is
often the focus of early stages of restoration. Large-scale
invasive species clearing programs are increasingly com-
mon, e.g. removal of Tamarix spp. from foodplains in
the southwestern US (Taylor & McDaniel 1998), mass
eradication of alien woody species from riparian areas in
South Africa (van Wilgen et al. 1998), and biocontrol of
Lythrum salicaria throughout the northern US (Blossey
et al. 2001)]. Large-scale restoration efforts often do not
plan for active revegetation. They assume that once the
barrier of the invasive species is removed, revegetation
will occur through natural recolonization (Galatowitsch
et al. 1999a). For example, natural recolonization is sug-
gested as the best strategy for revegetation in wetlands
following site preparation (Mitsch et al. 1998). However,
several studies have shown that failure to plant, results
in depauperate communities because of barriers to dis-
persal and establishment of native species (Lewis 1982;
Forbes 1999; Mulhouse & Galatowitsch 2003). Research
in California grasslands suggests native establishment
following invader removal is complicated by native
propagule availability and environmental modifcation
by the invader (Seabloom et al. 2003a, 2003b).
Despite the widespread application of clearing
programs and the debate on active revegetation, little is
known about how the transition from post control bare
ground to the establishment of native species occurs
(Kellogg & Bridgham 2002), save that initial dynamics
during this establishment phase are expected to have
effects on the ultimate plant community composition
(Weiner 1990). Studies of vegetation development fol-
lowing Pteridium aquilinum control in British heathlands
have shown that the direction and rate of vegetation
community establishment can be highly variable (Marrs
& Lowday 1992; LeDuc et al. 2000), and that vegetation