Bcl-2 and Bax Expression on Rat Ischemic Kidney
P. Eschwege, V. Paradis, M. Conti, S. Loric, F. Dumas, P. Berteau, M. Ahmed, S. Droupy,
b. Charpentier, A. Legrand, P. Bedossa, and G. Benoit
I
T has been suggested that oxidative stress can cause
injuries to the renal tubular epithelial cells.
1
Cells can
die by two distinct pathways: necrosis or apoptosis.
2
Necro-
sis is associated with metabolic collapse that leads to cell
swelling, loss of plasma membrane integrity, and cell rup-
ture. Apoptosis requires energy, and constitutes a strictly
regulated device responsible for the ordered removal of
superflous damaged cells. The family of Bcl-2–related pro-
teins constitutes one of the biologically most relevant
classes of apoptosis-regulatory gene products.
2,3
Bcl-2 as
Bax belongs to a family of apoptosis-regulatory gene prod-
ucts which may either be death antagonists (Bcl-2) or death
agonists (Bax). The relative amount of death agonists and
antagonists from Bcl-2 family constitutes a regulatory bal-
ance.
3
Anti-apoptotic properties of Bcl-2 oncoprotein are
associated with a marked decrease in the cellular genera-
tion of reactive oxygen species and prevention to lipid
peroxidation.
2
Moreover, Bcl-2 works by inhibiting the
mitochondrial generation of reactive oxygen species (ROS)
even under anaerobic conditions in which the generation of
ROS is reduced.
2
Apoptosis seems to play a key role in
acute tubular injury as it was described by Schumer et al.
4
after a brief period of ischemia and 24 to 48 hours after
reperfusion. Most of the studies described apoptosis pro-
cess after ischemia–reperfusion. Therefore, to our knowl-
egde, we do not know the pathway of postischemic cell
death: necrosis or apoptosis. To improve our knowlegde, we
have studied the occurrence of apoptotic process and the
expression of Bcl-2 and Bax proteins after warm renal
ischemia.
ANIMALS, MATERIALS, AND METHODS
Animals and Surgical Procedure
Thirty-six male Sprague-Dawley rats (body weight 350 to 400 g)
were studied. They were housed in metabolic cages during the
experiment with free access to feed and water. Anesthesia was
induced by intramuscular administration of ketamine (0.2 mg/100
g). A coeliotomy was performed, and the left and right renal
pedicles were gently dissected and clamped for 15, 30, 45, 60, and
120 minutes (n = 6 rats per group). One group was operated but
not clamped.
A right nephrectomy was performed after the end of the warm
ischemic period for immunohistochemical analysis. The right kid-
neys were collected, opened longitudinally and immediately im-
mersed in Bouin liquid until analysis.
Experimental Groups
The animals were divided into 6 groups: group 1 (n = 6),
unclamped control rats; group 2 (n = 6), renal pedicles clamped 15
minutes; group 3 (n = 6), renal pedicles clamped 30 minutes; group
4 (n = 6) renal pedicles clamped 45 minutes; group 5 (n = 6; renal
pedicles clamped 60 minutes; group 6 (n = 6), renal pedicles
clamped 120 minutes.
Immunohistochemical Method
A total of 36 Bouin-fixed paraffin-embedded kidney samples were
studied. Serial sections were performed for routine histologic
evaluation (HES) and for the immunohistochemical procedure.
Immunohistochemistry was performed using an automated immu-
nostainer (Techmate 500, Dako, Carpinteria, Calif) with the avi-
dine– biotin–peroxidase method. Anti-Bax rabbit polyclonal anti-
bodies 1/250 (Biogenex, San Ramon, Calif) and monoclonal mouse
anti Bcl-2 1/120 (Dako, Glostrup, Denmark) were used.
As positive controls, fixed paraffin-embedded rat spleen samples
were used. Localization and intensity of staining were indepen-
dently assessed by two pathologists who were not aware of the time
of warm ischemic periods. In case of discordance, a consensual
evaluation was performed by simultaneous reviewing of slides by
the two pathologists. The localization of immunostaining was
described and the intensity of staining was graded as absent (no
staining = 0) or present (+ to ++).
RESULTS
No significant morphologic lesion was observed in the
kidney tissue, whatever the duration of ischemia. Especially
we did not detect any apoptotic bodies. In positive control
experiments, several cells labeled with Bcl-2 and Bax anti-
bodies in the spleen tissue. No Bax and Bcl-2 immunostain-
From the Laboratoire de Chirurgie Expe ´ rimentale et Service
d’Urologie, Ho ˆ pital de Bice ˆ tre, Faculte ´ de Me ´ decine (P.E., M.A.,
S.D., G.B.) the Service d’Anatomie Pathologique, Ho ˆ pital de
Bice ˆ tre (V.P., P.B.) the Laboratoire de Biochimie, Ho ˆ pital de
Bice ˆ tre (M.C., A.L.), Le kremlin-Bice ˆ tre, France, the Laboratoire
de Biochimie A, Ho ˆ pital Necker, (S.L., F.D., P.B.), Paris, France
and the Service de Ne ´ phrologie, Ho ˆ pital de Bice ˆ tre (B.C.), Le
Kremlin-Bice ˆ tre, France.
Address reprint requests to Pascal Eschwege MD, Depart-
ment of Urology, Ho ˆ pital de Bice ˆ tre, 78 rue du Ge ´ ne ´ ral Leclerc,
94270 Le Kremlin-Bice ˆ tre, France.
Supported by grants from Fondation de L’Avenir etude ET-5
118, and Etablissement Franc ¸ ais des Greffes.
© 1998 by Elsevier Science Inc. 0041-1345/98/$19.00
655 Avenue of the Americas, New York, NY 10010 PII S0041-1345(98)00844-6
Transplantation Proceedings, 30, 2861–2862 (1998) 2861