180 Brain Research, 322 (1984) 180-183
Elsevicr
BRE 20459
In vivo chronic lead exposure alters pH]nitrendipine binding in rat striatum
R. A. RIUS, L. LUCCHI, S. GOVONI and M. TRABUCCHP
Institute of Pharmacology and Pharmacognosy, University of Milan, Milan and IChair of Toxicology,
II University of Rome, Rome (Italy)
(Accepted June 26th, 1984)
Key words: lead exposure - - calcium antagonist binding-- striatal membranes
The effect of lead as a neurotoxic agent has been associated with alterations in calcium metabolism. On this line, the present study
shows that lead alters the characteristics of [3H]nitrendipine ([3H]NDP) binding to rat striatal membranes. In vitro, lead shares the ac-
tion of calcium in enhancing [3H]NDP binding although it is more potent on a molar basis. In vivo, lead exposure through drinking wa-
ter enhances [3H]NDP binding to crude synaptosomal membrane preparations. This effect is lost when membranes are washed with
EDTA-EGTA, indicating that the increased binding is due to the persistence of lead in the brain of treated rats.
Lead poisoning is associated with a wide account of
symptoms pointing out brain impairment3,9.15; in fact,
lead intoxication modifies various neurochemical
parameters indicating an alteration of the functional
state of cholinergic 17,2°, noradrenergiO 22 GABAer-
gic 13,19 and dopaminergic neurotransmission 6. The
effect of lead on neurotransmitters may be at least
partially due to an interaction with calcium in the
mechanisms regulating release, uptake and metabo-
lism of neurotransmitters at neuronal level. In partic-
ular, in vitro data on striatal synaptosomes show that
lead potentiates calcium-dependent dopamine re-
lease. In addition, it has been shown that lead may
decrease the affinity of brain mitochondria for calci-
um 2,4, leading to an accumulation of free intracellular
calcium, which is harmful for the cell 23.
Calcium entry is a regulated process', in particular,
calcium channel opening (either voltage-dependent
or receptor-operated ~6) modulates the influx of calci-
um into the cell. The availability of labelled organic
calcium entry blockers made possible the study of
calcium channels in different tissues including the
brain ~2.1s. In the present investigation the effect of
lead on [3H]NDP binding was studied both by adding
this ion in vitro and after chronic in vivo exposure of
the animals through the drinking water.
Pregnant Sprague-Dawley rats (Charles River,
Calco, Italy) were used in our study. Animals at day
16 of pregnancy were caged individually and housed
at constant temperature (22 °C) and humidity
(50%). They were exposed to a light cycle of 12 h a
day (from 07.00 to 19.00 h) and had free access to
food and water. Mothers were given a lead acetate
solution (2.5 g/l; 1360 ppm lead) or sodium acetate as
control in their drinking water. The offspring were
reduced to 10 animals/litter after birth. At 21 days of
age the young rats were separated from their mothers
and divided by sex. They received the same drinking
solution which had been supplied to the mothers.
Lead-treated rats showed no significantly different
growing rate with respect to the age-matched con-
trois. At 6 weeks of age the animals were killed by de-
capitation. Brains were quickly removed, striatal tis-
sues dissected and frozen on dry-ice. Tissue was ho-
mogenized using an Ultra-Turrax homogenizer (set-
ting 2, 10 s) in 20 vols. of 0.32 M sucrose. The result-
ing homogenate was centrifuged at 1000 g for 10 min.
Supernatant was centrifuged again at 48,000 g for 15
min. The pellet was treated as follows: (A) resus-
pended in 25 vols. of 50 mM Tris-HCl pH 7.4 and
then centrifuged as before and used; or (B) exten-
sively washed with EDTA (10 mM)-EGTA (10 raM)
Correspondence: R. A. Rius, Institute of Pharmacology and Pharmacognosy, University of Milan, Via Andrea del Sarto 21, 20129
Milano, Italy.
0006-8993/84/$03.00 © 1984 Elsevier Science Publishers B.V.