Aviation, Space, and Environmental Medicine x Vol. 80, No. 5, Section II x May 2009 A15
Z WART SR, O LIVER SAM, F ESPERMAN JV, K ALA G, K RAUHS J, E RICSON
K, S MITH SM. Nutritional status assessment before, during, and after
long-duration head-down bed rest. Aviat Space Environ Med 2009;
80(5,Suppl.):A15–22.
Introduction: Bed rest is a valuable ground-based model for many of
the physiological changes that are associated with spaceflight. Nutri-
tional changes during and after 60 or 90 d of head-down bed rest were
evaluated. Methods: A total of 13 subjects (8 men, 5 women; ages 26–
54 yr) participated in either 60 or 90 d of bed rest. Blood and urine were
collected twice before bed rest and about once per month during bed
rest. Samples were stored frozen and batch analyzed. Data were ana-
lyzed using repeated-measures analysis of variance. Results: During bed
rest, markers of bone resorption (such as N-telopeptide excretion, P ,
0.001) increased and serum concentration of parathyroid hormone de-
creased ( P , 0.001). Also, oxidative damage markers such as superoxide
dismutase increased ( P , 0.05), and after 90 d of bed rest, total antioxi-
dant capacity decreased ( P , 0.05). During bed rest, iron status indices
showed patterns of increased iron stores with a decreased concentration
of transferrin receptors ( P , 0.01). Discussion: These changes are similar
to some of those observed during spaceflight, and further document the
utility of bed rest as a model of spaceflight.
Keywords: spaceflight, nutrition, weightlessness, bone resorption, para-
thyroid hormone, oxidative stress, antioxidants, iron, transferrin.
S
OME OF THE CLINICAL concerns for long-duration
spaceflight (longer than 30 d) are bone and muscle
loss, inadequate dietary intake, decreased nutrient stores,
increased oxidative damage due to factors such as ra-
diation exposure and increased iron stores, and altered
nutrient metabolism (41). Designing and testing counter-
measures to mitigate these negative physiological effects
of spaceflight require a solid understanding of changes in
nutrient status during spaceflight, because macro- and
micronutrients are essential for every cell and function in
the body. Data from 11 International Space Station (ISS)
astronauts suggest that their nutritional status is compro-
mised after long-duration spaceflight (40). Some of the
most striking changes are decrements in vitamin D, folate,
vitamin K, and vitamin E status, and increases in markers
of bone resorption and oxidative damage. Changes in uri-
nary excretion of phosphorus and magnesium were also
evident after 4 to 6 mo of spaceflight (40).
That vitamin D status is decreased after long-duration
spaceflight is clearly indicated by results of studies from
Skylab, Mir, and ISS missions (17,33,39,40). Even for
crewmembers who used supplements, 25-hydroxyvitamin
D status decreased (17,40). The low levels of vitamin D in
the space food supply and the absence of ultraviolet
light during spaceflight likely contribute to this phe-
nomenon. Altered vitamin D status is accompanied by
evidence of increased bone resorption during and after
spaceflight (31,33,40). During spaceflight, the excretion
rates of urinary markers of bone resorption are typically
100–150% of their preflight values (3,5,31,33,39). Loss of
bone mineral is increased by skeletal unloading during
weightlessness (10,21,33,36,39,41).
Evidence exists that during and after spaceflight, along
with changes in nutrient status, the metabolism of cer-
tain nutrients is altered (41), and iron is one of these nu-
trients. The data suggest that as a result of microgravity,
storage pools of iron are shifted so that less iron is in red
blood cells and more is stored in ferritin, and less iron is
transported (less transferrin) (35,40). Serum ferritin con-
centration is significantly increased, the amount of ferri-
tin iron is slightly greater than before launch, and the
amount of transferrin is decreased. Red blood cell mass
also decreases (1,6,14,43). After landing, a delay in re-
placing red blood cells often leads to decreased hemoglo-
bin, hematocrit, and mean corpuscular volume (34,40).
Like other physiological effects of weightlessness,
changes in nutrient status and metabolism can be stud-
ied in the bed rest analog of weightlessness. The qualita-
tive effects of bed rest on bone and calcium homeostasis
are similar to the effects of spaceflight, but the quantita-
tive effects are generally less than (about half) those of
spaceflight (41). As reviewed by Meck and colleagues
(23) and others (25), bed rest is a good model for space-
flight, but lack of standard procedures has limited the
ability to draw conclusions across studies. This report is
one of a series of reports on the Flight Analogs Project,
which is designed to lay the groundwork for a standard
bed rest protocol. Standard procedures were developed
From the Nutritional Biochemistry Laboratory, Human Adaptation
and Countermeasures Division, NASA Johnson Space Center, Houston,
TX; Enterprise Advisory Services, Inc., Houston, TX; Wyle, Houston,
TX; Indiana University-Purdue University, Fort Wayne, IN; and
Universities Space Research Association, Houston, TX.
Address reprint requests to: Scott M. Smith, Ph.D., NASA Johnson
Space Center, Mail Code SK, 2101 NASA Parkway, Houston, TX 77058;
scott.m.smith@nasa.gov.
Reprint & Copyright © by the Aerospace Medical Association,
Alexandria, VA.
DOI: 10.3357/ASEM.BR07.2009
Nutritional Status Assessment Before, During, and
After Long-Duration Head-Down Bed Rest
Sara R. Zwart, Susan A. Mathews Oliver,
J. Vernell Fesperman, Geeta Kala, Jane Krauhs,
Karen Ericson, and Scott M. Smith