MARINE RECORD Open Access
First record of Thysanozoon brocchii
(Platyhelminthes: Polycladida) from Indian
waters
Reshma Pitale
*
and Deepak Apte
Abstract
This work reports the occurrence of Thysanozoon brocchii from the rocky intertidal coast of Ratnagiri and Dwarka,
the West Coast of India, for the first time. Two morphotypes were found, the first morph has a buff brownish
papillate dorsal surface, with few specimens having white spots. The second morph has light coloured papillae
which form a distinct cross marking along the dorsal surface. The species possesses double male copulatory
apparatus with seminal vesicle, prostatic vesicle and sclerotized stylet. This cosmopolitan species has previously
been recorded from Italy and Mediterranean, Japan, South and West Africa, Florida, New Zealand, Brazil, and United
Kingdom. Synonymized species and older descriptions have been compared to examine similarities and
dissimilarities. Considering the existence of varied colour morphs of this species, a detailed comparative analysis of
morphological characters, reproductive histology and molecular framework is recommended.
Keywords: Pseudocerotidae, Colour patterns, Intraspecific variations, Reproductive anatomy, Cosmopolitan,
Maharashtra, Gujarat
Introduction
Polyclad flatworms are free living members of the
phylum Platyhelminthes. These coral reef and rocky
shore inhabitants are more diverse in the tropical envir-
onment (Prudhoe, 1985). Cryptic behavior, apparent
specificity with food preference, aposematic colouration
and mimicry with opisthobranch molluscs and fishes
(Ang & Newman 1998; Newman & Canon, 2005) make
them potentially significant in intertidal or reef ecology.
Members of the family Pseudocerotidae are peculiar
for their brilliant colour patterns. However, colour
variation is commonly seen forming species complex in
their systematics. Perhaps such complexities arising in
the taxonomy of these worms can be resolved by careful
observations of external as well as reproductive anatom-
ical features. Newman and Canon (1995) described three
species of the genus Pseudoceros from the Indo-Pacific
region showing remarkable similarity within external
appearance. They, too, emphasized the significance of
precise documentation of colour and patterns. Litvaitis,
et al. (2010) tested the coloration pattern of Pseudoceros
bicolor complex against molecular evidence and emerging
complexities in the taxonomy of the same.
Grube (1840) described the genus Thysanozoon con-
sidering the papillate dorsal surface, unlike the rest of
the Cotylean genera. Type species of the genus was
Thysanozoon diesingii, which has later been synony-
mized for T. brocchii. This genus includes about 23
species worldwide (Tyler, 2013). Laidlaw (1902) reported
Thysanozoon plehni from the Laccadive Island of India.
However, this species has been reviewed and assigned to
Acanthozoon plehni. After a gap of about a century, Apte
and Pitale (2011) mentioned a member of this genus
from Kavaratti, Lakshadweep Island, India. Later, Sreeraj
and Raghunathan (2013) reported Thysanozoon nigropa-
pilosum from South Andaman.
The present study reports Thysanozoon brocchii for
the first time from the Indian shoreline. Apart from
external morphology and reproductive anatomy, the
study encompassed the review of characteristic features of
some of the synonyms and a few previous descriptions,
and their comparisons with the two different morphs
presented herein.
* Correspondence: rd.pitale@bnhs.org
Bombay Natural History Society, Hornbill House, S.B. Singh Road, Mumbai,
Maharashtra 400 001, India
© The Author(s). 2017 Open Access This article is distributed under the terms of the Creative Commons Attribution 4.0
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reproduction in any medium, provided you give appropriate credit to the original author(s) and the source, provide a link to
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Pitale and Apte Marine Biodiversity Records (2017) 10:21
DOI 10.1186/s41200-017-0123-0