Sequence Variants of NAT1 and NAT2 and Other
Xenometabolic Genes and Risk of Lung and
Aerodigestive Tract Cancers in Central Europe
James D. McKay,
1
Mia Hashibe,
1
Rayjean J. Hung,
1,3
Jon Wakefield,
1
Valerie Gaborieau,
1
Neonila Szeszenia-Dabrowska,
4
David Zaridze,
5
Jolanta Lissowska,
6
Peter Rudnai,
7
EleonoraFabianova,
8
DanaMates,
9
LenkaForetova,
10
VladimirJanout,
11
VladimirBencko,
12
Amelie Chabrier,
1
Janet Hall,
1
Paolo Boffetta,
1
Federico Canzian,
2
and Paul Brennan
1
1
IARC, Lyon, France;
2
German Cancer Research Center-Deutsches Krebsforschungszentrum, Heidelberg, Germany;
3
University of
California at Berkeley, Berkeley, California;
4
Department of Epidemiology, Institute of Occupational Medicine, Lodz, Poland;
5
Institute of Carcinogenesis, Cancer Research Centre, Moscow, Russia;
6
Department of Cancer Epidemiology and Prevention,
Cancer Center and Maria Sklodowska-Curie Institute of Oncology, Warsaw, Poland;
7
National Institute of Environmental
Health, Budapest, Hungary;
8
Specialized Institute of Hygiene and Epidemiology, Banska Bystrica, Slovakia;
9
Institute
of Public Health, Bucharest, Romania;
10
Department of Cancer Epidemiology and Genetics, Masaryk Memorial
Cancer Institute, Brno, Czech Republic;
11
Department of Preventive Medicine, Faculty of Medicine, Palacky
University, Olomouc, Czech Republic; and
12
Charles University of Prague, First Faculty of Medicine,
Institute of Hygiene and Epidemiology, Prague, Czech Republic
Abstract
Tobaccosmokecontainsanextensivecocktailofhighly
carcinogenic chemicals. Individuals with a slower
elimination rate of the chemicals in tobacco smoke
may have increased exposure to their carcinogenic
properties compared with those with a faster rate.
Polymorphisms that alter the function of the genes
involved in the activation or the detoxification of the
chemical carcinogens in tobacco smoke can potentially
influence an individual’s risk of developing a tobacco-
related cancer. To test this hypothesis, we have
genotyped polymorphisms in 16 genes involved in
metabolism of chemical carcinogens in a Central and
Eastern European case-control study comprising 2,250
lung cases, 811 upper aerodigestive cancer (UADT)
cases, and 2,704 controls. The N-acetyltransferase
(NAT ) genes were the most implicated in risk, with
the NAT1*10 haplotypeshowinganinverseassociation
inlungcancer,inbothheterozygotecarriers[oddsratio
(OR), 0.81; 95% confidence interval (95% CI), 0.70-0.93]
and homozygote carriers (OR, 0.70; 95% CI, 0.48-1.01),
suggesting a genotype dose response (P < 0.001). In
UADT cancer, a similar inverse association was noted
in NAT1*10 although only in heterozygotes (OR, 0.78;
95%CI, 0.65-0.95). In NAT2 , when considering the
individuals inferred acetylator phenotypes based on
their NAT2 diplotype, ‘‘slow’’ acetylators compared
with intermediate or fast acetylators showed no
association with risk. None of the other 14 genes
provided robust evidence of an association for either
lung or UADT cancer. We therefore conclude that, of
the genetic variation studied, NAT1 genewasthemost
likely candidate to influence the risk of developing a
tobacco-related cancer. (Cancer Epidemiol Biomarkers
Prev 2008;17(1):141 – 7)
Introduction
Tobacco smoking is the predominant cause of lung and
upper aerodigestive tract (UADT) cancers (1). The
mutagenic chemical carcinogens in tobacco smoke are
eliminated from the human body via a series of enzymes
that metabolize the carcinogenic compounds. Genetic
variation in functionally important regions of xenometa-
bolic genes seem to influence the rate of action of these
enzymes (2-4). An individual’s genetic profile of xen-
ometabolic alleles will have a role in determining the rate
at which carcinogens are eliminated and therefore the
extent of carcinogen exposure. Individuals who have
inherited alleles that result in higher degree of carcinogen
exposure are expected to have a higher cancer risk relative
to those who inherited lower exposure alleles (2-6).
We hypothesized that polymorphisms in the genes
encoding the proteins involved in either the activation
(phase I) and/or the detoxification (phase II) of chemical
carcinogens in tobacco smoke will influence risk of
developing lung or UADT cancers. To test this hypothesis,
we have genotyped polymorphisms in 16 xenometabolic
genes in a large case-control study of lung cancer patients
and UADT cancer cases.
Materials and Methods
Study Participants. The study was conducted in
15 centers in six countries of Central and Eastern
Europe, including Czech Republic (Prague, Olomouc,
Brno), Hungary (Borsod, Heves, Szabolcs, Szolnok,
Cancer Epidemiol Biomarkers Prev 2008;17(1). January 2008
Received 6/18/07; revised 10/2/07; accepted 10/18/07.
Grant support: National Cancer Institute R01 grant (contract no. CA 092039-01A2)
and Association for International Cancer Research grant (contract no. 03-281).
The costs of publication of this article were defrayed in part by the payment of page
charges. This article must therefore be hereby marked advertisement in accordance
with 18 U.S.C. Section 1734 solely to indicate this fact.
Requests for reprints: James McKay, IARC, 150 cours Albert Thomas, F-69372 Lyon
Cedex 08, France. Phone: 33-4-727380993; Fax: 33-4-7273-8342. E-mail: mckay@iarc.fr
Copyright D 2008 American Association for Cancer Research.
doi:10.1158/1055-9965.EPI-07-0553
141
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