86 SURGERY
APOPTOSIS PROVIDES A PHYSIOLOGIC MEANS for the
clearance of tissue neutrophils (PMNs) and resolu-
tion of inflammatory processes.
1,2
Conversely, dys-
functional (delayed) PMN apoptosis has been
implicated in numerous hyperinflammatory dis-
ease states, including smoke inhalation injury,
3
the
acute respiratory distress syndrome,
4
and the sys-
temic inflammatory response syndrome.
5
Dysfunc-
tional PMN apoptosis may also be involved in the
pathogenesis of postinjury multiple organ failure
(MOF). We
6
and others
7-9
have reported that trau-
ma patients at risk for MOF have delayed PMN
apoptosis in the early postinjury period. Trauma
patients’ plasma delays the apoptosis of healthy
donors’ PMNs, indicating the presence of circulat-
ing antiapoptotic mediators.
6-8
We have recently
implicated platelet-activating factor (PAF) and
PAF-like phospholipids, including lyso-phos-
phatidylcholine (LPC) as the key circulating anti-
apoptotic mediators in trauma patients’ plasma.
10
Our previous epidemiologic studies identified
red blood cell (RBC) transfusion as the most robust
Clinically relevant hypertonicity prevents
stored blood- and lipid-mediated delayed
neutrophil apoptosis independent of p38
MAPK or caspase-3 activation
Walter L. Biffl, MD, Rachel Carnaggio, MS, Ernest E. Moore, MD, David J. Ciesla, MD,
Jeffrey L. Johnson, MD, and Christopher C. Silliman, MD, PhD, Providence, RI, and Denver, Colo
Background. Delayed apoptosis of primed neutrophils (PMNs) may facilitate PMN-mediated tissue
injury leading to postinjury multiple organ failure. Aged (42-day-old) stored red blood cells (RBC42)
delay PMN apoptosis through proinflammatory phospholipids such as platelet-activating factor (PAF)
and lyso-phosphatidylcholine (LPC). Hypertonic saline (HTS) attenuates PMN cytotoxic functions. We
hypothesized that clinically relevant HTS would provoke PMN apoptosis, as well as prevent stored blood-
and lipid-mediated delayed PMN apoptosis through activation of p38 mitogen-activated protein kinase
(MAPK) and caspase-3.
Methods. PMNs harvested from healthy volunteers were incubated (5% CO
2
, 37° C, 24 hr) with
RBC42 plasma, PAF (20 μM), or LPC (4.5 μM), with or without the p38 MAPK inhibitor SB
203580, the caspase-3 inhibitor zDEVD-fmk (10 μmol/L) or the pan-caspase inhibitor zVAD-fmk (20
μmol/L). Duplicate samples were preincubated in HTS (Na [180 mM]). Apoptotic index (% PMNs
undergoing apoptosis) was assessed morphologically. p38 MAPK activation was assessed by Western
blotting. Caspase-3 activity was measured colorimetrically.
Results. PAF, LPC, and RBC42 plasma delayed apoptosis; HTS increased apoptosis compared with con-
trols. HTS prevented PAF, LPC, and RBC42-delayed apoptosis. p38 MAPK was not activated by HTS;
its inhibition had no effect on the actions of HTS. Caspase inhibition attenuated the ability of HTS to
increase apoptosis, but it did not affect the ability of HTS to restore healthy PMN apoptosis in the pres-
ence of RBC42.
Conclusion. HTS increases PMN apoptosis and prevents stored blood- and lipid-mediated delayed PMN
apoptosis. HTS may activate caspase-3, but alternative signaling pathways appear to be involved in
modulating the effects of lipids on PMN apoptosis. (Surgery 2003;134:86-91.)
From the Departments of Surgery, Rhode Island Hospital/Brown Medical School, Providence, RI, and the Den-
ver Health Medical Center, and Bonfils Blood Center and Department of Pediatrics, University of Colorado
Health Sciences Center, Denver, Colo
Accepted for publication March 23, 2003.
Presented at the 1
st
Joint Meeting of the Surgical Infection Soci-
ety (22
nd
Annual Meeting) and Surgical Infection Society-
Europe (15
th
Annual Meeting), Madrid, Spain, May, 2002.
Supported in part by The Society of University Surgeons Foun-
dation Junior Faculty Fellowship Award (WLB), a National
Blood Foundation Scientific Research Grant (WLB), and
National Institutes of Health grants P50GM4922 and
T32GM08315 (EEM, WLB, CCS).
Reprint Requests: Walter L. Biffl, MD, Division of Trauma and
Surgical Critical Care, 593 Eddy Street, APC 110, Providence, RI
02903.
© 2003 Mosby, Inc. All rights reserved.
0039-6060/2003/$30.00 + 0
doi:10:1067/msy.2003.178