Research Article
A Computational Model for Investigating Tumor Apoptosis
Induced by Mesenchymal Stem Cell-Derived Secretome
Melisa Hendrata
1
and Janti Sudiono
2
1
Department of Mathematics, California State University, Los Angeles, CA 90032, USA
2
Department of Oral Pathology, Faculty of Dentistry, Trisakti University, Jakarta, Indonesia
Correspondence should be addressed to Melisa Hendrata; mhendra@calstatela.edu
Received 1 May 2016; Revised 1 August 2016; Accepted 8 August 2016
Academic Editor: Said Audi
Copyright © 2016 M. Hendrata and J. Sudiono. Tis is an open access article distributed under the Creative Commons Attribution
License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly
cited.
Apoptosis is a programmed cell death that occurs naturally in physiological and pathological conditions. Defective apoptosis can
trigger the development and progression of cancer. Experiments suggest the ability of secretome derived from mesenchymal stem
cells (MSC) to induce apoptosis in cancer cells. We develop a hybrid discrete-continuous multiscale model to further investigate the
efect of MSC-derived secretome in tumor growth. Te model encompasses three biological scales. At the molecular scale, a system
of ordinary diferential equations regulate the expression of proteins involved in apoptosis signaling pathways. At the cellular scale,
discrete equations control cellular migration, phenotypic switching, and proliferation. At the extracellular scale, a system of partial
diferential equations are employed to describe the dynamics of microenvironmental chemicals concentrations. Te simulation
is able to produce both avascular tumor growth rate and phenotypic patterns as observed in the experiments. In addition, we
obtain good quantitative agreements with the experimental data on the apoptosis of HeLa cancer cells treated with MSC-derived
secretome. We use this model to predict the growth of avascular tumor under various secretome concentrations over time.
1. Introduction
Apoptosis is a normal, genetically regulated process in which
a cell undergoes a sequence of intracellular complex processes
that trigger self-destruction. Cancers occur due to mutations
of certain fundamental genes that disable the cells to perform
apoptosis, giving rise to malignant tumor cells that grow
uncontrollably. With its genetic instability, an individual
tumor cell becomes a forerunner parent cell that has the
potential to develop into a cluster, biologically complex tumor
consisting of approximately 10
6
cells.
Various cancer treatments have been explored with the
ultimate goal of suppressing its growth and spreading and
perhaps even eradicating cancerous cells. Recently, mes-
enchymal stem cells (MSCs) have become a topic of great
focus in relation to cancer. MSCs are known to secrete a broad
panel of proteins including growth factors, chemokines, and
cytokines, which are called secretome [1]. Growing evidence
suggests that MSCs have an important role in afecting the
behavior of tumor cells [2]. While some studies reported
that MSCs favor tumor growth, others showed that MSCs
can suppress tumorigenesis [3, 4]. In particular, it has been
reported that secretome contained in conditioned media
(CM) of MSCs promotes apoptosis and autophagy of cancer
cells [5]. Experiments done by Sandra et al. show that
secretome signifcantly induces apoptosis in HeLa cancer
cells in concentration and time dependent manner [6].
From intracellular perspective, there are two well-known
major signaling pathways leading to apoptosis: the intrinsic
pathway centered on mitochondria and the extrinsic pathway
initiated by death receptors called Tumor Necrosis Factor
(TNF). Tere is now evidence showing that these two path-
ways are connected and afect one another [7, 8]. Moreover,
recent research has also revealed the third pathway, called the
perforin pathway, which involves T-cell mediated cytotoxicity
and is induced by granzyme B protein. Perforin pathway is
also connected to the intrinsic pathway and all three pathways
eventually converge into the activation of caspase 3 protein
leading to cell death, chromatin condensation, chromosome
Hindawi Publishing Corporation
Computational and Mathematical Methods in Medicine
Volume 2016, Article ID 4910603, 17 pages
http://dx.doi.org/10.1155/2016/4910603