RESEARCH PAPER
Limnology (2007) 8:121–130 © The Japanese Society of Limnology 2007
DOI 10.1007/s10201-007-0202-8
Barbara Leoni · Giuseppe Morabito · Michela Rogora
Davide Pollastro · Rosario Mosello · Silvia Arisci
Elena Forasacco · Letizia Garibaldi
Response of planktonic communities to calcium hydroxide addition in a
hardwater eutrophic lake: results from a mesocosm experiment
Abstract The main focus of this study was to investigate the
effects of lime treatment with Ca(OH)
2
on a hypereutrophic
hardwater lake (Lake Alserio, Northern Italy), focusing on
its impact on plankton communities. We performed a me-
socosm experiment using two large enclosures, one for
treatment and one control, in the lake. The addition of
Ca(OH)
2
proved to be effective in reducing phosphorus
concentration, but the effect was limited in time. Redissolu-
tion of the precipitating calcium carbonate as it came into
contact with the deep water determined a phosphorus in-
crease about 2 weeks after the liming. The liming had a
strong impact on the plankton assemblage, as was demon-
strated by the sharp decrease in phytoplankton and zoo-
plankton biomass and density, and 1 month after the
treatment we observed a different taxonomic composition
inside the enclosure from that of the lake. The result of the
experiment showed that liming with Ca(OH)
2
can be re-
garded as a suitable treatment for accelerating recovery of
the lake. Multiple treatments will be necessary to improve
the trophic level over a longer period.
Key words Eutrophication · Restoration · Liming Ca(OH)
2
·
Phytoplankton · Zooplankton
Introduction
Eutrophication as a result of anthropogenic impact is one
of the commonest causes of the deterioration of the quality
of water bodies located in the most populated and industri-
alized areas of the world. Since the end of the 1960s, many
ecotechnological methods have been proposed and used to
tackle the negative effects of the eutrophication processes
(see Straskraba 1996 for a review). Management at water-
shed level is essential to reduce nutrient loading to the lake
so that further interventions within the water body can be
effective.
Most of these recovery actions are primarily designed to
reduce the amount of phosphorus available to the phyto-
plankton, by adding to the lake water substances that
determine the precipitation of phosphorus through its in-
corporation into insoluble compounds. This results can be
achieved using aluminium or ferric compounds; however,
both treatments may have negative side effects, such as the
aluminium poisoning of aquatic organisms in the absence
of neutralizing agents, or an unacceptable pH decrease in
low buffered systems (Cooke 1993).
In the last few years, liming with CaO, Ca(OH)
2
, and
CaCO
3
has been successfully used to treat natural ponds
(Sosnowska 1990), man-made ponds (Murphy et al. 1990),
experimental enclosures (Koschel 1997), and hardwater
lakes (Prepas et al. 2001; Zhang 2001). In hardwater lakes,
the addition of lime simulates the natural process of calcite
biogenic precipitation, also known as “whiting.” This proc-
ess, which usually occurs during periods of high productiv-
ity, is caused by the precipitation of calcium carbonate
when the lake water is supersaturated with it. The situation
is favored by increasing pH resulting from the CO
2
consumption by phytoplankton. During calcium carbonate
precipitation, phosphorus is incorporated into the calcite
crystals and precipitates out of solution.
Insofar as it imitates a natural process, liming has fewer
negative effects on biota than other chemicals used to con-
trol algal biomass. Moreover, lime addition can also induce
phytoplankton flocculation and decrease water transparen-
cy, further reducing phytoplankton biomass and slowing
down the photosynthetic activity.
There are few studies describing the effect of liming on
freshwater biotic communities, mostly restricted to acidic
lakes (Olem 1991), where the planktonic assemblages are
quite different from those commonly found in hardwater
eutrophic lakes. Some information on planktonic response
to liming in hardwater lakes comes from the experiments
B. Leoni (*) · E. Forasacco · L. Garibaldi
Dipartimento di Scienze dell’Ambiente e del Territorio, Università
degli Studi di Milano Bicocca, Piazza della Scienza 1, 20126 Milano,
Italy
Tel. +39 0264482737; Fax +39 0264482795
e-mail: barbara.leoni@unimib.it
G. Morabito · M. Rogora · D. Pollastro · R. Mosello · S. Arisci
CNR Istituto per lo Studio degli Ecosistemi, Verbania Pallanza, Italy
Received: October 10, 2006 / Accepted: January 24, 2007 / Published online: July 7, 2007