SHORT REPORT
ABSTRACT: The surface EMG area often exhibits progressive enlarge-
ment during a submaximal fatiguing contraction, but the underlying reasons
still remain uncertain. Fatigue-induced changes in the surface-detected mo-
tor unit action potentials (S-MUAPs) of 10 human thenar motor units (MUs)
with widely differing physiological properties were examined. After 2 min of
repetitive 40-Hz stimulation, the size of the S-MUAPs of all MUs increased,
the magnitude of which was negatively correlated with their tetanic tension
changes. These findings suggest that during muscle fatigue, in addition to
reflecting recruitment of new MUs and increases in firing rates of the active
MUs, the surface EMG may also be markedly influenced by changes in the
S-MUAPs, especially in fast fatigable muscles.
© 1998 John Wiley & Sons, Inc. Muscle Nerve 21: 1786–1789, 1998
DISSOCIATION OF THE ELECTRICAL
AND CONTRACTILE PROPERTIES IN
SINGLE HUMAN MOTOR UNITS
DURING FATIGUE
K. MING CHAN, MD, FRCP(C),
1
LEONARD P. ANDRES, MSc,
2
YELENA POLYKOVSKAYA, MD,
1
and WILLIAM F. BROWN, MD, FRCP(C)
1
1
Department of Neurology, New England Medical Center,
Boston, Massachusetts, USA
2
Nutrition, Exercise Physiology & Sarcopenia Laboratory, Jean Mayer USDA
Human Nutrition Research Center on Aging at Tufts University,
Boston, Massachusetts, USA
Accepted 5 May 1998
Surface EMG is commonly used in the study of
muscle fatigue. The progressive enlargement in its
rectified integrated area during submaximal fatigu-
ing contraction is thought to primarily reflect an in-
crease in the number of recruited motor units
(MUs) and in their firing rates.
7,10,14
A third possi-
bility is that the size of the motor unit action poten-
tial (MUAP) might change during fatiguing contrac-
tion. Such possibility indeed has been shown in
animal studies,
1,8,18
but has not been systematically
investigated in humans. Therefore, the purposes of
this study were to examine: (1) the effect of fatigue
on the size of the MUAPs; and (2) the relationship of
the latter changes to the contractile characteristics of
these MUs.
METHODS
We recently described a noninvasive method for
studying the contractile and electrical properties of
human thenar MUs.
2,3
To briefly summarize, single
motor axons with the lowest stimulation threshold
were excited by finely graded electrical impulses de-
livered through a surface bipolar stimulator placed
at multiple points over the accessible portions of the
median nerve at the wrist and upper arm. Their as-
sociated MUAPs were detected by using self-adhesive
surface recording electrodes (1.5 × 1 cm) with the
active electrode placed over the thenar muscles and
the reference electrode over the first metacarpopha-
langeal joint. The tension output of each motor unit
was detected by a Grass FT-10 force transducer
placed against the first interphalangeal joint aligned
in the plane of its prime vector. Using this method,
the twitch properties, tetanic tensions, and electrical
properties of 10 MUs from 5 young healthy subjects
(1–3 MUs per subject) have been thoroughly char-
acterized and reported.
3
The fatigue response of
each MU was determined by delivering 13 stimuli at
40 Hz every second for a total of 2 min.
1
Changes in
the size of the surface-detected MUAP (S-MUAP)
Abbreviations: EMG, electromyography; FI, fatigue index; MU, motor
unit; MUAP, motor unit action potential; S-MUAP, surface-detected motor
unit action potential
Key words: human; muscle fatigue; motor unit action potential; potentia-
tion; Na/K pump
Correspondence to: Dr. K. Ming Chan, Division of Neuroscience, 513
Heritage Medical Research Center, University of Alberta, Edmonton, Al-
berta, T6G 2S2 Canada
CCC 0148-639X/98/121786-04
© 1998 John Wiley & Sons, Inc.
1786 Short Reports MUSCLE & NERVE December 1998