ELSEVIER
Marine Chemistry 48 (1994) 71-82
Time courses of intracellular and extracellular lipid classes in batch
cultures of the toxic dinoflagellate, Gymnodinium cf. nagasakiense
Christopher C. Parrish a, Guy Bodennec b, Patrick Gentien b
aOcean Sciences Centre and Department of Chemistry, Memorial University of Newfoundland, St. John's, NF, AI C 5S7, Canada
bD~partement Environnement Littoral, IFREMER, Centre de Brest, 29280 PlouzanO. France
Received 16 July 1993; revision accepted 27 April 1994
Abstract
The effect of temperature and light on lipid production was determined for an isolate of Gymnodinium cf. nagasak&nse
which is known to be toxic. Samples were taken in a time course in cultures grown at 13 or 18°C and at 35 or 75
#E.m -2 .s -~ . The lipid class composition of Gymnodinium cf. nagasakiense cells and their exudates were analyzed with
the Chromarod-Iatroscan system. The main intracellular lipid class, glycolipid, was present at similar concentrations in
the cells in all three cultures. Maxima in intracellular concentrations of triacylglycerol, and minima in intracellular free
fatty acid concentrations occurred near the time of maximum culture density. Dissolved free fatty acid concentrations
were higher during the first 20 days than in the following 10 days of each culture. The culture grown at 13°C had the
highest intracellular triacylglycerol concentration and the highest extracellular free fatty acid concentration. The culture
grown at 35 #E.m -2- s-l had the highest extracellular glycolipid concentration. Gas chromatographic analyses at the end
of the growth experiments showed that pentaenoic fatty acids consisting mainly of 18 : 5n3 were twice as prominent in
cells grown at the lower light level. However, the amount of 18 : 5n3 released into the surrounding medium in this culture
was negligible.
1. Introduction
In recent years there has been a global increase in
the number of documented toxic algal blooms. This
is partly a result of greater awareness of such events
in the scientific community, but it is also likely a
consequence of increased coastal pollution and use
of coastal waters for aquaculture purposes
(Anderson, 1989). Toxic events appear to have
expanded in geographic extent and variety. In
eastern Canada, for example, while the St.
Lawrence estuary and the Bay of Fundy have
long been recognized as areas of paralytic shellfish
poisoning, the first recorded case of such poisoning
occurred in Newfoundland in 1982 (White and
White, 1985). In 1987, a very unusual diatom
bloom was responsible for the toxicity of mussels
in neighbouring Prince Edward Island (Bates et al.,
1989).
Some phytoplankton are thought to cause fish
kills by producing lipid compounds containing
polyunsaturated fatty acids (Yasumoto et al.,
1990). Such compounds may also be toxic to
other algae, thus permitting the toxic alga to fully
utilize available nutrients for growth (Gentien and
Arzul, 1990a,b) resulting in a catastrophic bloom.
By modifying their environment these algae may
thus proliferate where they would not otherwise
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