INTRODUCTION
For the last decade, there has been a steady increase in the glob-
al acreage of transgenic commercial crops. In 1998 it was esti-
mated that there were over 28 million ha of transgenic com-
mercial crops grown worldwide, excluding China (James, 1998).
Because honey bees (Apis mellifera) play critical roles in pollinat-
ing fruits and vegetables (Morse & Calderone, 2001), many stud-
ies have been conducted to evaluate the effect of transgenic
products on honey bees (reviewed by Malone & Pham-Delègue
et al. 2001).
The majority of studies on the effects of transgenic crops on
honey bees have used purified transgene products, i.e. proteins.
There have been five laboratory studies done on purified Bacil-
lus thuringiensis (Bt) proteins (Sims, 1995; Arpaia, 1996; Malone
et al., 1999; US Environmental Protection Agency, 2000; Malone
et al., 2001). Nine studies tested the effects of serine protease
inhibitors on honey bees (Belzunces et al., 1994; Malone et al.,
1995; Burgess et al., 1996; Malone et al., 1998; Girard et al., 1998;
Jouanin et al., 1998; Malone et al., 1999; Pham-Delègue et al.,
2000; Malone et al., 2001). The effects of other proteins such as
chitinase, ß-1,3 glucanase and avidin have been tested in labora-
tory studies (Picard-Nizou et al., 1997; Malone et al., 2002).
There have been only two published studies that used intact
pollen instead of purified transgene proteins (Jouanin et al., 1998;
Schur et al., 2000; cited in Malone & Pham-Delègue, 2001).
While it is important to determine the potential detrimental
effects of transgene products on honey bees, it is also important
to determine whether pollen from transgenic plants has nega-
tive effects on honey bees. This is because there might be other
new proteins expressed in the transgenic plants besides the
transgene product(s), due to either pleiotropic effects of the
inserted genes (Uberlacker et al., 1996), alternative splicing
(Lopato et al., 1999), or other mechanisms such as interactions
between host genes and inserted genes (Kohli et al., 1998).
Because some Bt toxins (such as Cry1A(b) and Cry1F) target
lepidopteran insects and honey bees store large amounts of
pollen in the hive (Winston, 1987), we suspect that Bt corn
pollen could be an alternative control method against the greater
wax moth (Galleria mellonella). The greater wax moth is an
important pest of honey bees. Larvae of the greater wax moth
cause considerable damage to beeswax combs left unattended
by bees. Beeswax combs in weak or dead colonies and those
placed in storage are subject to attack (Caron, 1999). Although
there are other methods available for controlling this pest
(Cantwell, 1981), Bt corn pollen, if effective at killing wax moth,
would be economical and unlikely to contaminate honey or
beeswax as do current fumigant control methods.
The objectives of this study were two-fold: (1) to determine the
effect of Bt corn pollen on honey bee worker larvae, and (2) to
evaluate Bt corn pollen as a potential control of the greater wax
moth.
MATERIALS AND METHODS
Honey bee pollen-feeding studies
Corn pollen was collected during corn tasseling in late July or
early August (2000 and 2001), depending on the variety of corn,
from commercial farms (regular corn or Cry1A(b) corn), or
Michigan State University (USA) experimental farms (Cry1F
corn). Corn tassels were bagged (Lawson No. 402 - Shower-
proof’d
®
) in the early morning or in the afternoon when tassels
began to shed pollen. On the following day, the bags were invert-
ed, tapped to release the pollen and removed. After the bags
*Corresponding author: pett@msu.edu
Journal of Apicultural Research 42(4): 77–81 (2003) © IBRA 2003
ORIGINAL ARTICLE
Effects of dietary transgenic Bt corn pollen
on larvae of Apis mellifera and Galleria
mellonella
ANNE V HANLEY ,
1
ZACHARY Y HUANG
1,2
AND WALTER L PETT
1
*
1
Department of Entomology, Michigan State University, East Lansing, MI 48824, USA
2
Ecology, Evolutionary Biology and Behavior Program, Michigan State University, East Lansing, MI 48824, USA
Received 4 April 2003, accepted subject to revision 13 August 2002, accepted for publication 21 August 2003
SUMMARY
The effects of dietary transgenic Bt corn pollen on 4–5-day-old honey bee worker larvae were examined. We
measured larval and pupal mortalities, pupal weight, and haemolymph protein concentration of newly emerged
adults after they were fed (as larvae) various pollens (mixed bee pollen, non-transgenic corn pollen, Cry1A(b)
or Cry1F corn pollen). There were no significant differences in all the parameters tested between larvae fed
transgenic Bt corn pollen Cry1A(b) or Cry1F and non-transgenic corn pollen. We also evaluated Bt corn
pollen as a potential control for wax moth larvae in a laboratory study. We fed first instar wax moth larvae
three types of pollen: non-transgenic corn pollen, Cry1A(b) corn pollen, and Cry1F corn pollen. We found that
the mortality of larvae fed Cry1F corn pollen was significantly greater than the mortality of larvae fed Cry1A(b)
corn pollen or non-transgenic corn pollen (P < 0.05). In each trial Cry1F fed larvae showed 100% mortality.
Our studies suggest that transgenic Bt corn pollen does not pose a threat to honey bee larval development and
has the potential to serve as an alternative control for wax moth.
Keywords: Bacillus thuringiensis, Bt corn pollen, greater wax moth, Galleria mellonella, honey bees Apis mellifera,
transgenic pollen