JANS Journal of Applied and Natural Science 6 (1): 315-319 (2014)
Ultramorphology of antennal sensilla of open-nesting honey bees Apis florea
F. and Apis dorsata F. (Hymenoptera: Apidae)
Neelima R. Kumar, Kalpna Nayyar, Ruchi Sharma and Anudeep*
Department of Zoology, Panjab University, Chandigarh-160014, INDIA
*Corresponding author. E- mail: anuresearch_aedes@yahoo.com
Received: March 3, 2014; Revised received: May 30, 2014; Accepted: June 20, 2014
Abstract: Antenna of honey bees is geniculate and has been known to possess a wide variety of sensilla. The
sensilla are sensitive to odor, temperature, humidity, air pressure and gustatory stimuli. In the present investigation,
ultrastructural study on the antenna of the two open-nesting worker honey bees- Apis florea F. - the dwarf honey
bee and A. dorsata F. - the giant honey showed considerable amount of variation in the types and distribution
pattern of sensilla in the two species. The antennal form as well as the sensilla arrangement has been suggested to
be adapted to the pheromone perception need of regarding a particular species.
Keywords: Apis florea, Apis dorsata, Antenna, SEM, Sensilla
INTRODUCTION
South-East Asia is the centre of honey bee diversity
and the evolutionary homeland of honey bees. Apis
florea and A. dorsata are open-nesting bees which
build single comb. The head appendages constitute an
important morphological feature to study honey bee
systematics. The antenna of honey bees is important
for understanding behavior of honey bees. It consist
of a basal scape, pivoted pedicel and a ten-segmented
flagellum. Antennae are important for sensory
perception. Various types of sensilla are observed on
the segments of flagellum. According to
Wigglesworth (1965), the antennae are the main sites
of olfactory receptors in most insects. Six types of
antennal sensilla have been observed on the flagellum
of honey bee A. mellifera drone and worker. They
have been referred to as a thick basiconic sensillum, a
tapered basiconic sensillum, a trichoid sensillum, a
placoid sensillum, a coeloconic or ampullaceal
sensillum and a coelocapitular sensillum, formerly
known as campaniform sensillum (Kuwabara and
Takeda, 1956; Dietz and Humphreys, 1971). SEM
studies on antennal sensilla of honey bee workers
have been performed by Al-Ghamdi (2006) from
Saudi Arabia. Suwannapong and Wongsiri (2004)
described occurrence of sensilla trichodea types A, B,
C and D on different segments of flagellum from
Thailand. Therefore, the aim of the present study is to
document the diversity of Apis species by using the
scanning electron microscopic technique of an impor-
tant morphological character i.e. the antenna.
ISSN : 0974-9411 (Print), 2231-5209 (Online) All Rights Reserved © Applied and Natural Science Foundation www.ansfoundation.org
MATERIALS AND METHODS
Study material: Two open-nesting species- Apis
florea F. and A. dorsata F. were taken for the present
study. A. dorsata and A. florea are found wild in
nature. A. dorsata contributes towards large amount
of forest honey and bees wax harvested by tribal peo-
ple of India.
Study area and sample collection: A. florea was
collected from nests located in a hedges and bushes in
Panjab University Campus while A. dorsata was
collected while foraging on flowers in Botanical
garden, Panjab University, Chandigarh.
Preservation: The collected material of Apis florea
and Apis dorsata was preserved in 70% alcohol and
the protocol of Bozolla and Russell (1999) was
followed.
Preparation of material for scanning electron
microscopy: The antennae were carefully excised
from the freshly collected worker bees of Apis florea
and Apis dorsata. These were then washed with
phosphate buffer. The samples were fixed in 5%
gluteraldehyde for 2 hrs. Subsequently these were
washed with phosphate buffer 2 to 3 times and then
dehydrated through graded series of acetone and dried
in a critical point drier. Dehydrated samples were
mounted on slides in the desired orientation with the
help of double side adhesive tape under binocular
microscope. The samples were attached in such a
way that they became visible from all sides. The stubs
were placed inside the sputter for gold coating to
overcome the problem of “charging” and “beam dam-
age”. The sputtered specimens were examined in Jeol
2008
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