American Journal of Plant Sciences, 2014, 5, 3652-3661
Published Online November 2014 in SciRes. http://www.scirp.org/journal/ajps
http://dx.doi.org/10.4236/ajps.2014.524381
How to cite this paper: Handayani, M., Sasaki, H., Matsuda, R., Takechi, K., Takano, H. and Takio, S. (2014) Characterization
of an Epiphytic Bacterium Neptunomonas sp. BPy-1 on the Gametophytes of a Red Alga Pyropia yezoensis. American Jour-
nal of Plant Sciences, 5, 3652-3661. http://dx.doi.org/10.4236/ajps.2014.524381
Characterization of an Epiphytic Bacterium
Neptunomonas sp. BPy-1 on the
Gametophytes of a Red Alga Pyropia
yezoensis
Midia Handayani
1
, Hiroyuki Sasaki
1
, Ryuya Matsuda
1
, Katsuaki Takechi
1
,
Hiroyoshi Takano
1
, Susumu Takio
1,2
1
Graduate School of Science and Technology, Kumamoto University, Kumamoto, Japan
2
Center for Marine Environment Studies, Kumamoto University, Kumamoto, Japan
Email: stakio@gpo.kumamoto-u.ac.jp
Received 3 October 2014; revised 2 November 2014; accepted 16 November 2014
Copyright © 2014 by authors and Scientific Research Publishing Inc.
This work is licensed under the Creative Commons Attribution International License (CC BY).
http://creativecommons.org/licenses/by/4.0/
Abstract
The established culture of gametophytes of the red alga Pyropia yezoensis (TU-1) is superficially
colonized by epiphytic bacteria. By 16S rRNA sequencing, 6 bacterial species were identified, and a
culturable bacterium, Neptunomonas sp. BPy-1, was isolated. The 16S rRNA sequences of BPy-1
showed 100% identity with that of Neptunomonas sp. 0536, a probiotic bacterium found in green-
shell mussels in New Zealand. Physiological tests revealed that 22 characters were identical be-
tween BPy-1 and 0536, but that 4 characters differed. BPy-1 cannot grow in the artificial seawater
used for the culture of gametophytes. BPy-1 can grow in the artificial seawater with ethanol or
butanol but not in methanol or propanol. To determine the effect of BPy-1 on gametophyte growth,
BPy-1 colonization was reduced by 80% using a multi-enzyme cleaner. Changing the cleaner con-
centration yielded two types of gametophytes, a compressed or callus-like form and a nearly nor-
mal form. BPy-1 promoted the growth of the treated gametophytes with relatively normal form,
while it showed less effect on compressed gametophytes. These findings suggested that BPy-1
promotes the growth of damaged gametophytes but does not affect the development of normal
gametophytes.
Keywords
Neptunomonas, Pyropia yezoensis, Ethanol-Eating Bacteria, Epiphytic Bacteria