Molecular Ecology Notes (2007) 7, 191–193 doi: 10.1111/j.1471-8286.2006.01587.x
© 2006 The Authors
Journal compilation © 2006 Blackwell Publishing Ltd
Blackwell Publishing Ltd
TECHNICAL ARTICLE
A fast and inexpensive DNA extraction/purification protocol
for brown macroalgae
GALICE HOARAU, JAMES A. COYER, WYTZE T. STAM and JEANINE L. OLSEN
Department of Marine Benthic Ecology and Evolution, Centre for Ecological and Evolutionary Studies, University of Groningen,
PO Box 14, 9750 AA Haren, The Netherlands
Abstract
Here we describe a rapid method for extracting DNA from dried brown algae material
using a microtitre plate system in conjunction with a milling instrument. The method
allows the preparation of nuclear and organelle DNA of quality suitable for polymerase
chain reaction amplification. It combines high throughput with low cost per sample: DNA
from 192 samples can be extracted in c. 3 h for < $0.40 per sample, nearly tenfold cheaper
than commercially available kits. Furthermore, by using microtitre plates, efficient storage
and downstream processing is facilitated.
Keywords: brown algae, DNA extraction, heterokonts, high throughput, Phaeophyceae
Received 15 June 2006; revision accepted 18 September 2006
Extracting polymerase chain reaction (PCR)-amplifiable
DNA from brown algae is notoriously difficult as poly-
saccharide and polyphenolic compounds, which can be
strong enzyme inhibitors, are often co-extracted (Mayes
et al. 1992; Jin et al. 1997). Protocols involving extraction
with cetyltrimethyl ammonium bromide (CTAB), precipit-
ation with chloroform-isoamyl alcohol and purification
with either gradient-centrifugation or glass-binding (e.g.
Sephaglas, Amersham Biosciences) techniques, have been
developed to overcome the problems (Coyer et al. 1994,
2002b; Phillips et al. 2001). While these methods yield PCR-
quality DNA, they are time-consuming and cannot be
automated, thereby precluding their usage in high throughput
projects. Methods employing Chelex resin are quick and
inexpensive and have been successfully used in some
brown and red algal species (Billot et al. 1998; Cohen et al.
2004), but not in other genera of brown algae (e.g. Fucus,
Turbinaria, Undaria; G. Hoarau, unpublished data). Further-
more, as DNA extractions with Chelex resin have a limited
shelf life, extractions must be repeated regularly (e.g.
Laminaria, Billot 1999; fish, G. Hoarau, unpublished data).
The limited shelf life and species-specific success therefore
preclude routine use of Chelex resin for DNA extraction
in brown algae. Alternatively, several commercial DNA
extraction kits for plants are available (e.g. Nucleospin
plant kit, Clontech; DNeasy 96 Plant Mini Kit, QIAGEN)
and have been successfully used with brown algae (Engel
et al. 2003; Billard et al. 2005). DNA extraction with kits is
much faster and can be automated, but the cost is often
prohibitive for large-scale projects. The increased costs
associated with greater samples sizes is an important
consideration when funding sources become limited. In
view of these problems, we developed a DNA extraction/
precipitation/purification method for brown algae that,
when combined with a mixer mill for pulverizing dried
samples, is fast, automated, and inexpensive.
Reagents included:
1 The CTAB extraction buffer was 2% CTAB, 1.4 m NaCl,
20 mm EDTA (pH 8), 100 mm Tris-HCl (pH 8), 0.1%
polyvinylpolypyrrolidone (PVPP), 0.2% β-mercaptoethanol
(added freshly) (Coyer et al. 1994).
2 The binding buffer consisted of 6 m NaI and 0.1 m Na
2
SO
3
,
clarified by filtration through a 0.45-μm membrane filter
(Whatman).
3 Silica fines were prepared by placing 20–30 g of silica
(silica gel 60 0.015–0.040, Merck) into c. 500 mL of milliQ-
filtered de-ionized water and stirring for c. 1 h. After
stirring, the silica was allowed to settle for c. 15 min. The
supernatant (= fines) was transferred to 50 mL plastic
tubes and centrifuged for 5 min at 1250 g. Note that the
silica that settled by gravity when preparing the fines
can still be used for other DNA extraction protocols
Correspondence: Dr Galice Hoarau, Fax: +31 503632261; E-mail:
g.g.hoarau@rug.nl