Human Reproduction vol.8 no. 1 pp.11 —16, 1993
Immunohistochemical localization of acidic and basic
fibroblast growth factors in normal human endometrium
and endometriosis and the detection of their mRNA by
polymerase chain reaction
R.A.Ferriani
1
, D.S.Charnock-Jones
1
, A.Prentice
1
,
E.J.Thomas
2
and S.K.Smith
13
Department of Obstetrics and Gynaecology, 'University of
Cambridge and
2
University of Southampton, Southampton, Hants,
UK
3
To whom correspondence should be addressed at: Department of
Obstetrics and Gynaecology, University of Cambridge, The Rosie
Maternity Hospital, Cambridge CB2 2SW, UK
Growth factors play a role in the cyclical growth and
vascularization of normal endometrium. Abnormal
endometrial proliferation and neovascularization may result
in endometriosis. This study determines the presence and
localization of acidic and basic fibroblast growth factors
(aFGF and bFGF respectively) in endometrium of normal
women, and in normal and ectopic endometrium of women
with endometriosis. Endometrium was obtained at curettage
or hysterectomy for benign disease, or laparoscopy for
endometriosis. aFGF- and bFGF-immunoreactivity was
detected at different phases of the menstrual cycle by
immunohistochemistry using primary polyclonal rabbit
antibodies. Expression of mRNA for aFGF and bFGF was
determined in normal endometrium by nested reverse trans-
criptase polymerase chain reaction (RT-PCR). aFGF- and
bFGF-immunoreactivity were both detected in endometrium
from normal women, and in normal and ectopic endometrium
of women with endometriosis. The pattern of staining with
the two different FGFs was the same: immunoreactivity was
predominantly confined to glandular epithelial cells and did
not change throughout the menstrual cycle. Little or only light
staining was seen in stromal cells and myometrium, and the
pattern of staining did not differ between endometriotic and
normal tissue. The presence of mRNA for aFGF and bFGF
was demonstrated in normal endometrium. The detection of
aFGF and bFGF mRNA in normal endometrium and aFGF-
and bFGF-immunoreactivity in normal and endometriotic
tissues suggests that these peptides may play a role in the
proliferation and angiogenesis of normal and ectopic human
endometrium.
Key words: endometrium/endometriosis/growth factors/immuno-
histochemistry/polymerase chain reaction
Introduction
Proliferation and repair of human endometrium after menstru-
ation is a prerequisite for implantation and development of the
fertilized ovum. To complete this function, endometrium
undergoes cyclical proliferation and differentiation of both
epithelial and stromal cells and neovascularization (Noyes et al.,
1950; Weitlauf, 1988). The paracrine and autocrine factors
regulating this growth are poorly understood, although these
modifications were assumed to be related to cyclical changes in
circulatory levels of ovarian steroids. Recently, diverse growth
factors have been associated with regulatory mechanisms of the
female reproductive tract, and growth factors may mediate the
oestrogen-stimulated growth response (Presta, 1988; Haining
et al., 1991a,b; Irwin etal, 1991; Nelson etal, 1991).
Fibroblast growth factors (FGFs) are a family of at least seven
structurally related peptides with high affinity for heparin
(Gospodarowicz et al., 1987; Rifkin and Moscatelli, 1989). The
most intensively studied members of the family are acidic FGF
(aFGF) and basic FGF (bFGF), which have 55% sequence
homology and a similar spectrum of biological activity, with
bFGF being 30 times more potent than aFGF (Gospodarowicz
et al., 1987). Both growth factors promote mitogenesis and
differentiation of a wide variety of neuroectodermal and meso-
dermal cells, and are associated with haemostasis, angiogenesis,
tissue repair, embryonic development and tumorigenesis
(Gospodarowicz etal., 1987; Rifkin and Moscatelli, 1989).
bFGF is present in a number of cultured cell types, normal tissues
and tumours (Gospodarowicz et al., 1987; Folkman et al., 1988;
Cordon-Cardo et al., 1990; Gonzalez et al., 1990) and its mRNA
has been cloned from human cDNA libraries (Abraham et al.,
1986).
Evidence that bFGF may be involved in endometrial function
includes the identification of angiogenic molecules in rabbit
endometrium (Lynch et al., 1986), and the identification of bFGF
in porcine (Brigstock et al., 1990) and human uterus (Cordon-
Cardo et al., 1990; Rusnati et al., 1990). Recombinant bFGF
stimulates plasminogen activator production in normal endo-
metrium and endometrial adenocarcinoma cell lines (Presta, 1988;
Rusnati et al., 1990) and stimulates proliferation of human
endometrial stromal cells (Irwin et al., 1991). Ovarian steroid
hormones modulate the synthesis and function of bFGF in
endometrial cells (Presta, 1988; Irwin et al., 1991), suggesting
that these growth factors may play a role during the cyclical
growth and vascularization of endometrium. Abnormal endo-
metrial proliferation and neovascularization within the peritoneal
cavity may result in endometriosis (Smith, 1991) and bFGF has
not previously been described in endometriosis.
There are fewer studies of aFGF. aFGF is less well conserved
between different species, and Southern blotting analysis of
human genomic DNA shows one gene for each growth factor
(Mergia et al., 1986). aFGF is found predominantly in brain and
retina (Bohlen et al., 1985), whilst capillary endothelial cells that
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