© by PSP Volume 23 – No 8. 2014 Fresenius Environmental Bulletin
1852
EFFECT OF SOLAR RADIATION,
TEMPERATURE AND SALINITY ON THE SURVIVAL
OF TWO DIFFERENT STRAINS OF ESCHERICHIA COLI
Slaven Jozić
1,
*, Mira Morović
1
, Mladen Šolić
1
, Nada Krstulović
1
and Marin Ordulj
2
1
Institute of Oceanography and Fisheries, Split, Croatia
2
Univesity of Split, University Department of Marine Studies, Split, Croatia
ABSTRACT
The simultaneous effect of temperature, salinity and
solar radiation, as well as the history and strain of bacte-
rial cells on Escherichia coli (E. coli) survival in seawater
under experimental and natural conditions were studied.
The experiments were carried out within the natural range of
temperature (12
o
C, 18
o
C and 24
o
C) and salinity (30.0 psu
and 36.5 psu). Natural samples of microbiologically con-
taminated sea water were taken during September 2011,
when the temperature and salinity of sea water were sta-
ble (23–24
o
C, 36–37 psu). In the absence of solar radia-
tion, the mean T
90
values differed, depending on the bac-
terial strain and were 42.50 h for E. coli ATCC 35218 and
33.55 h for E. coli ATCC 8739. No significant effect of
temperature or salinity on T
90
was found, but a strong and
significant negative effect of solar radiation on T
90
of both
E. coli strains was recorded. Depending on the bacterial
strain, the dominant effect of solar radiation reduced the
T
90
of E. coli by 15- to 70-fold. Within the ultraviolet A
(UVA) and photosynthetically active radiation (PAR) spec-
trum of solar radiation, the wavelengths of 320–360 were
found to be most bactericidal. If exposed to solar radia-
tion, sea water samples were found to be depleted of cul-
turable E. coli cells even during 24 h storage under ap-
propriate conditions. A higher resistance of wild E. coli
cells to the negative effects of environmental conditions
than cultivated cells was also found.
KEYWORDS: Escherichia coli, bacterial strain, survival, tempera-
ture, salinity, solar radiation
1. INTRODUCTION
When indicator organisms enter the marine environ-
ment, they experience a very hostile environment. The
hostile impact of the marine environment to these organ-
isms is reflected in the negative effects of the complex
* Corresponding author
array of physical, chemical and biological factors of the
marine environment, with a dominant effect of tempera-
ture, salinity and solar radiation [1-4].
The distribution and number of indicator bacteria de-
pends mostly on their input, but also on environmental
factors [5] and adaptation capacity. The adaptation capac-
ity of microbial cells to marine conditions is very minor,
which leads to physiological injury that might be sub-
lethal [6, 7] or lethal [8, 9]. If exposed to the negative
effect of some abiotic factors, particularly to solar radia-
tion, indicator bacteria rapidly enter into a temporary state
in which they lose culturability on standard bacteriologi-
cal media, but can still maintain some metabolic activity
[10, 11], infective capacity and potential for pathogenicity
[12, 13]. If further exposed to an unfavourable marine
environment, bacterial cells are irreversibly damaged and
die. For how long they will maintain culturability and sur-
vive in sea water, depends on many factors, such as the
intensity of environmental factors and the characteristics of
the bacterial cell, where besides the origin and pre-
exposure history, the bacterial strain plays an important
role [4, 14, 15]. Although there are many studies in which
the effects of the aforementioned parameters have been
addressed, their simultaneous effects have been poorly
investigated. This study is the first report of the separate
and simultaneous effect of temperature, salinity and solar
radiation, as well as the history and strain of E. coli cells
on their survival in sea water.
2. MATERIALS AND METHODS
2.1 Experiments with pure bacterial cultures
Experiments were performed with two different E. coli
strains that showed completely different morphological
characteristics on standard Tryptone bile X-glucuronide
(TBX) plates: E. coli ATCC 35218 originating from canine
faeces and E. coli ATCC 8739 originating from human
faeces. Bacterial suspensions for experiments were ob-
tained from pure cultures of test microorganisms incu-
bated on mineral-modified glutamate broth (MMGB). Bac-