PCR/RFLP Assay and Fungi 21
MOLECULAR BIOTECHNOLOGY Volume 31, 2005
RESEARCH
21
Molecular Biotechnology © 2005 Humana Press Inc. All rights of any nature whatsoever reserved. 1073–6085/2005/31:1/021–028/$30.00
*Author to whom all correspondence and reprint requests should be addressed. National Risk Management Research Laboratory, U.S.
Environmental Protection Agency, 109 T.W. Alexander Drive, Research Triangle Park, NC 27711. E-mail: dean.timothy@epa.gov.
Abstract
A Simple Polymerase Chain Reaction/Restriction Fragment Length
Polymorphism Assay Capable of Identifying Medically Relevant
Filamentous Fungi
Timothy R. Dean,* Michael Kohan, Doris Betancourt, and Marc Y. Menetrez
Because of the accumulating evidence that suggests that numerous unhealthy conditions in the indoor
environment are the result of abnormal growth of the filamentous fungi (mold) in and on building surfaces,
it is necessary to accurately reflect the organisms responsible for these maladies and to identify them in
precise and timely manner. To this end, we have developed a method that is cost effective, easy to perform,
and accurate. We performed a simple polymerase chain reaction/restriction fragment length polymorphism
(PCR/RFLP) analysis on multiple members of species known to negatively influence the indoor environ-
ment. The genera analyzed were Stachybotrys, Penicillium, Aspergillus, and Cladosporium. Each organism
underwent PCR with universal primers that amplified ribosomal sequences generating products from 550 to
600 bp followed by enzymatic digestion with EcoRI, HaeIII, MspI, and HinfI. Our results show that using
this combination of restriction enzymes enables the identification of these fungal organisms at the species
level.
Index Entries: PCR; RFLP; Stachybotrys chartarum; fungal identification; filamentous fungi.
1. Introduction
The identification, control, and removal of
filamentous fungi (mold) from the indoor envi-
ronment has become an important area of re-
search in recent years. This is because unwanted
growth and dissemination of fungal spores and
mycelial fragments negatively contribute to the
quality of indoor air. It has been estimated that
around 40% of all the homes in North America
contain mold growth, whereas in Northern Eu-
rope the proportion is between 20 and 40% (1,2).
Of the mold species that have been identified in
homes, currently only a small fraction have actu-
ally been implicated in adverse health effects. It
is probable that with increased awareness and
additional research the number of fungi that con-
tribute to an unhealthy indoor environment will
increase.
Fungal genera such as Stachybotrys, Penicil-
lium, Aspergillus, and Cladosporium are organisms
that have been isolated from unhealthy buildings
and are potentially associated with negative health
effects in humans (3,4). These health effects may
include itchy eyes, stuffy nose, headache, fatigue,
and in severe cases idiopathic pulmonary hemosi-
derosis (IPH) in infants (5–11). Necessity dictates
that to provide building owners, building occupants,
and remediators accurate information, methods of
fungal screening and identification be developed
that are rapid, unambiguous, highly specific, and
cost effective.
From a historic perspective, fungal isolates have
traditionally been identified based on macroscopic
and microscopic examination, analyzing the dis-
tinctions present in their colonies such as color and
texture along with conidial size, shape, and struc-