Research Article
Diagnosis of the Diatom Community upon Biofilm Development
on Stainless Steels in Natural Freshwater
Caroline Richard,
1
Smita Mitbavkar,
2
and Jessem Landoulsi
3,4
1
Laboratoire de M´ ecanique et Rh´ eologie (LMR), EA 2640, Polytech’Tours, Universit´ e Franc ¸ois Rabelais de Tours,
37200 Tours, France
2
CSIR-National Institute of Oceanography, Dona Paula, Goa 403 004, India
3
Sorbonne Universit´ es, UPMC Univ Paris 06, 75005 Paris, France
4
CNRS, UMR 7197, Laboratoire de R´ eactivit´ e de Surface, 75005 Paris, France
Correspondence should be addressed to Caroline Richard; caroline.richard@univ-tours.fr
and Jessem Landoulsi; jessem.landoulsi@upmc.fr
Received 26 January 2017; Accepted 13 April 2017; Published 25 May 2017
Academic Editor: David Alsteens
Copyright © 2017 Caroline Richard et al. Tis is an open access article distributed under the Creative Commons Attribution
License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly
cited.
Tis paper reports the development of bioflms on stainless steels (SS) upon exposure in a natural freshwater ecosystem for about
six months and focuses on the composition of diatom populations. By using environmental scanning electron microscopy (ESEM)
technique, we provide a detailed description regarding diatom identifcation at species level as well as their main characteristics,
including type, morphology, ability to form colony, and motility. Results reveal the presence of both prostrate (initial colonizers)
and stalked (late colonizers) forms. Pennate diatoms, Cocconeis placentula and Amphora cofeaeformis, and a centric diatom,
Melosira varians, are shown to be the abundant forms regardless of the SS type. Pennate diatoms dominate the community and
are directly attached to the substratum, whereas the centric form is entangled in the bioflm matrix in a signifcant number. Te
dominance of adnate forms suggests that these cells are sturdy and successfully maintaining their population. In situ monitoring of
the electrochemical response of immersed materials showed ennoblement of the open circuit potential, which seems to be due to
the biogenic production of H
2
O
2
, detected in a signifcant amount within the bioflms. Te substantial enrichment of bioflms with
diatoms potentially suggests the implication of these microorganisms in the process of ennoblement. A mechanism is proposed in
this paper describing the possible interactions of diatom community with SS in the studied ecosystem.
1. Introduction
Diatoms are microscopic, unicellular, autotrophic algae
which are members of the heterokontic group of cells
and are responsible for a major portion of the primary
productivity, thus forming the base of the food chain in
aquatic ecosystems. Based on their morphology, two types of
diatoms are identifed, centric with a radial symmetry and
pennate with a bilateral symmetry. In general, the centric
types are abundant in the water column, whereas the pennate
ones are found in bioflms formed on natural or artifcial
surfaces. Tis is due to their ability to attach to substrata via
extracellular polymeric substances (EPS) which are secreted
through a structure known as a raphe [1]. Depending on
the positioning and number of raphes, pennate diatoms are
classifed as monoraphid (one raphe), biraphid (two raphes,
both present on one valve or one on each valve), and araphid
(no raphe). Some diatom species attach to substrata with the
help of structures such as stalks or pads [2]. Both centric
and pennate types are ofen observed in each other’s habitats.
As centric types do not possess raphes, they are found
entangled within the bioflm community. Both forms occur
as single cells or as colonies in various shapes (flaments,
ribbons, fans, zigzags, or even stellate). Within the bioflms,
these diatoms play a similar role as their counterparts in
the water column forming the base of the benthic food
webs. Although the role of diatoms as primary producers in
aquatic environments is of major importance, their presence
on artifcial man-made structures is a serious issue. Diatom
fouling may, indeed, greatly afect the performance of metallic
Hindawi
Scanning
Volume 2017, Article ID 5052646, 13 pages
https://doi.org/10.1155/2017/5052646