Figen Esin Kayhan et al./ Elixir Pollution 110 (2017) 48560-48563 48560
The 2,4-D Herbicidal Effect of Defense Enzyme Activities and AChE
Levels in Liver and Gill Tissues of Xiphophorus hellerii (Swordtail fish)
Figen Esin Kayhan
*
, Gamze Tuğçe Turan, Güllü Kaymak, Cansu Akbulut and Nazan Deniz Yön Ertuğ
Marmara University, Science and Letters Faculty Department of Biology, İstanbul, Turkey.
Introduction
Fish are vulnerable to alteration of environmental factors
caused by the introduction of industrial wastes, heavy metals,
pesticides, oils, pills and other agents that directly affect the
aquatic ecology. Toxic effects of pesticides on the
biochemical and physiological systems of a fish can be
determined through the study of malondialdehyde levels
(MDA) and acetylcholine esterase enzyme activity (AChE),
among other methods [1]. Biochemical biomarkers can
provide information about the process of pesticide
detoxification.
When in contact with an organism, the toxic agent can
be biotransformed by enzymes, which act to make the
xenobiotic substance a less toxic compound and facilitate its
excretion. Lipid peroxidation may be the first step of cellular
membrane damage and can be induced by environmental
pollutants as herbicides. Organisms have both enzymatic and
non-enzymatic antioxidant defenses against reactive oxygen
species (ROS) [2].
One of the first enzymes that act in defense against ROS
is superoxide dismutase (SOD). It catalyses the conversion of
reactive superoxide anions (O
2
.-
) to hydrogen peroxide
(H
2
O
2
), which is subsequently detoxified by catalase (CAT)
and glutathione dependent peroxidase (GPx). GPx catalyses
the metabolism of H
2
O
2
to water involving a concomitant
oxidation of reduced glutathione (GSH), one of the most
important non-enzymatic antioxidants in the cell [3]. The
activity of AChE is another biochemical biomarker normally
used to monitor aquatic environments mainly contaminated
by pesticides. This enzyme can be inhibited by different
types of agrochemicals [4].
Pesticide pollution can be observed in increasing
amounts in water and soil as a result of its uncontrolled and
unaware usage in many countries. These pesticides can be
removed by rain and irrigation water. The number of
scientific studies show that the agricultural usage of
pesticides have negative ecological effects and toxic effects
on living things in the medium, environment in addition to
the effects on immune systems and various tissues, which
respectively increases day by day [5-8].
Two groups of herbicides can be divided in terms of
creation of oxidative stress: The first is known to be directly
responsible for the enhancement of free radical generation,
entering redox cycles and constantly generating ROS. The
second group of herbicides consists of several classes of
compounds known to be mainly inhibitors of antioxidant
enzymes, such as SOD and CAT [2].
2,4-Dichlorophenoxyacetic acid (2,4-D) has been the
most commonly used acidic phenoxy herbicide in agriculture
and forestry since 1940. 2,4-D is one of the most common
and most toxic herbicides. 2,4-D has a notorious past. It was
one of the two chemicals in the defoliants Agent Orange and
Agent Purple. It was also one of the two chemicals in Agent
White. The toxicity of 2,4-D depends on its chemical forms,
including salts, esters, and an acid form. The ester forms of
2,4-D can be highly toxic to fish and other aquatic life. 2,4-D
generally has moderate toxicity to birds and mammals, is
sorely toxic to fish and aquatic invertebrates, Herbicidal
activity of 2,4-D may also be due to an increase in the
production of ROS. The objective of this study is to
understand the effects of the herbicide 2,4-D on swordtail
fish tissues by using a set of biochemical markers. To this
aim, CAT and AChE enzyme activities and MDA levels were
measured using spectrophotometric methods in liver and gill
tissues of Swordtail fish (Xiphophorus hellerii Heckel, 1848).
Material and Method
The applied doses of 2,4-D were prepared using
commercial 2,4-Dichlorophenoxyacetic acid salt emulsions.
X. hellerii was obtained from local commercial aquarists. The
animals were acclimatized for two weeks in stock tanks
under laboratory conditions. Millipore water was used in this
experiment. During the acclimatization period, fish were fed
ad libitum with pellet twice a day. The fish were randomly
selected and divided into four experimental groups, arranged
Tele: 9002163464553
E-mail address: figenesink@gmail.com
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ARTICLE INFO
Article history:
Received: 12 July 2017;
Received in revised form:
19 September 2017;
Accepted: 29 September 2017;
Keywords
Oxidative stress in fish,
Herbicide,
AChE,
Antioxidant systems.
Elixir Pollution 110 (2017) 48560-48563
Pollution
Available online at www.elixirpublishers.com (Elixir International Journal)
ABSTRACT
In the present study, herbicide 2,4-D was investigated for its potency to induce oxidative
stress and its effects on antioxidant systems in gill and liver tissues of ( Xiphophorus
hellerii). Animals were exposed to sublethal doses of 2,4-D for 96 hours except for the
control group. Protein, malondialdehyde, catalase and acetylcholine esterase enzyme
activity (AChE) were determined using spectrophotometric methods. The results show
that protein levels were reduced in all experiments when compared to control group.
Levels of malondialdehyde were increased in each group. Catalase enzyme activity was
significantly decreased in all groups. In addition, an increased AChE activity was
observed.
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