Plant Cell, 1issue and Organ Culture 44: 155-159, 1996. 155
© 1996 KluwerAcademic Publishers. Printed in the Netherlands.
Growth and regenerability of long-term suspension cultures of the U.S. rice
cultivar Mercury
Herry S. Utomo, Timothy P. Croughan & Suzan S. Croughan*
Rice Research Station, Louisiana State University Agricultural Center, P.O Box 1429, Crowley, LA 70527-1429,
U.S.A. (* requests for offprints)
Received29 December1994;accepted in revisedform9 November1995
Key words: Oryza sativa L., regeneration, tissue culture, rejuvenation
Abstract
Suspension cultures of the U.S. rice cultivar Mercury have been maintained in modified General Medium for more
than 3 years. These suspensions have continued to have high and relatively stable regeneration rates. Two different
explants, immature panicles and seeds, were compared during the development of these embryogenic suspensions.
Initial formation of secondary embryogenic callus from immature panicles on induction medium was greater than
that from seeds. Suspensions of these two cell lines, however, did not differ morphologically and maintained
similar regeneration rates. After 5 months in culture the rates of regeneration began to decline. The suspensions
were plated onto regeneration medium without growth regulators for 2 weeks and then embryogenic cells were
manually selected and used to develop secondary suspensions. Through this simple rejuvenation procedure, the
suspensions retained high and stable regeneration rates. Variability in suspension growth, however, was observed
during the culture period. Slower growth occurred at weeks 13, 15, 27, and 29 and was associated with a decrease
in regeneration rates. Reproductive fertility of regenerated plants remained high for 3.5 years but then declined.
Abbreviations: CH- casein (acid hydrolysate); 2,4-D- 2,4- dichlorophenoxyacetic acid; MS - Murashige & Skoog
basal medium; SE- standard error
Introduction
The use of suspension cells of rice (Oryza sativa L.)
can facilitate the accomplishment of several research
objectives. They have been utilized to generate trans-
formed rice plants, through both particle bombardment
and polyethylene glycol treatment (Cao et al., 1992;
Lee et al., 1991), and for in vitro screening (Yamada et
al., 1986). Rice suspension cells also have been wide-
ly utilized as a source for protoplasts that are used for
somatic hybridization (Kyozuka et al., 1989), the pro-
duction of protoclones (Kanda et al., 1988), transfor-
mation (Zhang & Wu 1988; Li etal., 1992; Shimamoto
et al., 1989; Toriyama et al., 1988; Zhang et al., 1988;
Davey et al., 1991), and in vitro screening (Xie, 1991;
Utomo, 1994). In other species, suspension cells have
been used for rapid and continuous production of uni-
form biomass for studies of basic plant cell physiolo-
gy (Fry, 1990), enzymology (Herzbeck & Hiisemann,
1985), and secondary metabolites (Hall et al., 1988).
Establishment, selection, and preferential culture
methodology of embryogenic callus have been recog-
nized as critical factors in maintaining long-term mor-
phogenetic potential in cultures (Vasil & Vasil, 1984).
In recent years, the establishment of embryogenic cul-
tures has been achieved with several rice genotypes
previously regarded as recalcitrant (Lee et al., 1989;
Datta et al., 1990; Datta et al., 1992). This resulted
from better recognition of the conditions required to
produce embryogenic calluses (Vasil & Vasil, 1991;
Datta et al., 1992). However, characterization and
maintenance of long-term embryogenic rice suspen-
sions has not been reported. This paper describes the
characteristics of embryogenic suspension cells of the
U.S. rice cultivar Mercury that have been in culture for
over 3 years.